Asplenium bradleyi

D. C. Eaton

Bull. Torrey Bot. Club 4: 11. 1873.

Common names: Bradley's spleenwort
Endemic
Synonyms: Asplenium stotleri Wherry
Treatment appears in FNA Volume 2.

Roots not proliferous. Stems short-creeping to ascending, occasionally branched; scales dark reddish to brown throughout, narrowly deltate, (2–) 3–5 × 0.2–0.4 mm, margins entire or shallowly dentate. Leaves monomorphic. Petiole reddish or purplish brown throughout, lustrous, 1–10 (–13) cm, 1/3–3/4 length of blade; indument of brown, narrowly lanceolate scales at base, grading into hairs. Blade narrowly oblong to lanceolate, pinnate-pinnatifid to 2-pinnate, 2–17 (–20) × 1–6 cm, thin to moderately thick, sparsely pubescent; base truncate or obtuse; apex acute, not rooting. Rachis reddish or purplish brown proximally, fading to green in distal 1/3–2/3, lustrous, sparsely pubescent. Pinnae in (3–) 5–15 (–25) pairs, ovate, obovate to lanceolate or deltate-lanceolate; medial pinnae 6–40 × 3–10 mm; base truncate to obliquely obtuse; margins dentate to denticulate; apex acute or rounded. Veins free, barely evident. Sori 3 to numerous pairs per pinna, on both basiscopic and acroscopic sides. Spores 64 per sporangium. 2n = 144.


Habitat: Acidic rocks, usually on steep ledges
Elevation: 0–1000 m

Distribution

V2 326-distribution-map.gif

Ala., Ark., Ga., Ill., Ky., Md., Mo., N.J., N.Y., N.C., Ohio, Okla., Pa., S.C., Tenn., Va., W.Va.

Discussion

Asplenium bradleyi is a morphologically variable species, the allotetraploid derivative of A. montanum × platyneuron (W. H. Wagner Jr. 1954; D. M. Smith and D. A. Levin 1963; C. R. Werth et al. 1985). The sterile diploid form of A. bradleyi has been collected twice in nature (W. H. Wagner Jr. et al. 1973; A. M. Evans 1988), and isozyme studies indicate that the allotetraploid has had a polytopic origin (C. R. Werth et al. 1985b). Occurring rarely to locally in the Appalachian region, A. bradleyi overlaps with both progenitor taxa, but it is fairly frequent in the Ozark and Ouachita region where A. montanum is absent. Sterile hybrids with A. pinnatifidum (A. × gravesii), A. montanum (A. × wherryi D. M. Smith et al.), and A. platyneuron are known from nature.

Selected References

None.

Lower Taxa

None.

... more about "Asplenium bradleyi"
not rooting +
Rounded (?) +  and Acute (?) +
Warren H. Wagner Jr. +, Robbin C. Moran +  and Charles R. Werth +
D. C. Eaton +
truncate +  and obliquely obtuse +
simple +  and 4-pinnate +
17 cm170 mm <br />0.17 m <br /> (20 cm200 mm <br />0.2 m <br />) +
2 cm20 mm <br />0.02 m <br /> (17 cm170 mm <br />0.17 m <br />) +
narrowly oblong;lanceolate pinnate-pinnatifid +
thin;moderately thick +
Bradley's spleenwort +
Ala. +, Ark. +, Ga. +, Ill. +, Ky. +, Md. +, Mo. +, N.J. +, N.Y. +, N.C. +, Ohio +, Okla. +, Pa. +, S.C. +, Tenn. +, Va. +  and W.Va. +
0–1000 m +
Acidic rocks, usually on steep ledges +
monomorphic +
fertile +  and than sterile ones +
dentate +  and entire +
dentate +  and denticulate +
0.6 cm6 mm <br />0.006 m <br /> (4 cm40 mm <br />0.04 m <br />) +
0.3 cm3 mm <br />0.003 m <br /> (1 cm10 mm <br />0.01 m <br />) +
perforate +, reticulate +, spiny +  and winged +
not articulate +
10 cm100 mm <br />0.1 m <br /> (13 cm130 mm <br />0.13 m <br />) +
purplish brown +  and reddish +
1/3 length of blade;3/4 length of blade +
c--shaped +
1 cm10 mm <br />0.01 m <br /> (10 cm100 mm <br />0.1 m <br />) +
Bull. Torrey Bot. Club +
fading +  and green +
not proliferous +
0.2 cm2 mm <br />0.002 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
brown +  and dark reddish +
0.3 cm3 mm <br />0.003 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
lanceolate +  and deltate +
0.2mm;0.4mm +
acroscopic +  and basiscopic +
less lunate;linear +
dorsiventral +  and symmetric +
short-creeping +  and ascending +
Asplenium stotleri +
Asplenium bradleyi +
Asplenium +
species +
anastomosing +