Asteraceae (tribe Heliantheae) subtribe Clappiinae

H. Robinson

Phytologia 41: 39. 1978.

Treatment appears in FNA Volume 21. Treatment on page 251. Mentioned on page 5, 220, 253.

Subshrubs or shrubs, 5–40 cm (often succulent or fleshy). Leaves mostly cauline; opposite (proximal) or distally or mostly alternate; sessile; blades mostly linear to filiform (terete or subterete), margins entire, faces glabrous. Heads radiate, borne singly. Calyculi 0. Involucres hemispheric or obconic. Phyllaries persistent, 12–16 in 2–4+ series (distinct, obovate or ovate to lanceolate or linear, unequal, striate with resinous nerves in Clappia). Receptacles convex, usually epaleate (paleae rarely scattered, subulate scales; receptacles sometimes with setiform or subulate enations). Ray-florets 2–11 or 10–15, pistillate, fertile; corollas yellowish or yellow. Disc-florets 20–100+, bisexual, fertile; corollas yellowish or yellow, tubes shorter than to longer than funnelform throats, lobes 5, deltate to narrowly lanceovate; anther thecae pale; stigmatic papillae in 2 lines. Cypselae clavate to columnar (10–12-ribbed, sparsely hairy); pappi persistent, either of 12–25 subulate scales in 1 series, or of ca. 50 bristles or setiform scales in 3–4 series.

Distribution

sw United States, Mexico

Discussion

Genera 2, species 3 (2 genera, 3 species in the flora).

Traditionally, Clappia has been included in Helenieae and Pseudoclappia in Senecioneae. H. Robinson (1981) suggested that the two genera are related to Varilla (Varillinae).

Selected References

None.

Lower Taxa

Key

1 Ray florets 10–15; pappi of 12–25 subulate scales in 1 series Clappia
1 Ray florets 2–4; pappi of ca. 50 bristles or setiform scales in 3–4 series Pseudoclappia
not tailed +, rounded +  and obtuse +
connate +  and distinct +
subequal +
scarious +
usually ovate +  and lanceolate +
usually deltate +  and lanceolate +
spiciform +, racemiform +, paniculiform +  and corymbiform +
indeterminate +  and determinate +
Theodore M. Barkley† +, Luc Brouillet +  and John L. Strother +
H. Robinson +
decurrent +
compound +  and simple +
mostly linear;filiform +
winged;ribbed;winged;ribbed +
tuberculate +  and rugose +
barbellulate +  and barbellate +
yellow +  and yellowish +
clavate +  and columnar +
fertile +  and bisexual +
fertile +  and bisexual +
sw United States +  and Mexico +
pistillate +, staminate +  and neuter +
winged +  and beaked +
dispersed +
indeterminate +
homogamous +  and heterogamous +
each +  and sessile +
obconic;hemispheric +
alternate +, or +  and opposite +
deltate +  and narrowly lanceovate +
2-carpellate +
persistent +
12 +  and 16 +
bearing subulate enations +, hairy +  and bristly +
12 +  and 25 +
setiform +  and subulate +
exalbuminous +
prostrate +  and ascending +
appendaged +  and truncate +
hirsutulous +  and glabrous +
papillate +
Compositae +
Asteraceae (tribe Heliantheae) subtribe Clappiinae +
Asteraceae tribe Heliantheae +
subtribe +
shorter than to longer than funnelform throats +
perennial +, biennial +  and annual +
shrub +  and subshrub +