Asteraceae (tribe Heliantheae) subtribe Marshalliinae

H. Robinson

Phytologia 41: 42. 1978.

Treatment appears in FNA Volume 21. Treatment on page 456. Mentioned on page 7.

Perennials, (10–) 20–80 (–120) cm. Leaves mostly basal and/or cauline; alternate; petiolate or sessile; blades 1-nerved or 3-nerved, mostly elliptic, linear, oblanceolate, ovate, or spatulate, margins entire, faces usually glabrous, glanddotted. Heads discoid, borne singly or in open, corymbiform arrays. Calyculi 0. Involucres turbinate to obconic or hemispheric. Phyllaries persistent, 10–20 in 2+ series (distinct, narrowly oblong to linear-elliptic or lance-subulate, subequal, herbaceous, basally thickened-keeled, sometimes callous or scarious-margined). Receptacles convex to conic, paleate (paleae mostly linear to filiform, herbaceous). Ray-florets 0. Disc-florets (20–) 45–90 (–160+), bisexual, fertile; corollas lavender, pink, purple, or white, tubes much longer than to equaling funnelform throats, lobes 5, lance-linear to linear-oblong; anther thecae cream to purple; stigmatic papillae in 2 lines. Cypselae stoutly obpyramidal, 5 (–6) -angled, hairy; pappi persistent, of 5 (–6) lance-deltate to lance-attenuate, scarious (often hyaline) scales.

Distribution

c, e United States

Discussion

Genus 1, species 7 (7 species in the flora).

Marshalliinae was related to epaleate subtribes of Heliantheae by H. Robinson (1981), rather than to Eupatorieae, as suggested by T. F. Stuessy (1977[1978]) and by B. L. Turner and A. M. Powell (1977[1978]). B. G. Baldwin and B. L. Wessa (2000) suggested that Marshallia may be sister to Gaillardiinae and results presented by Baldwin et al. (2002) showed Marshallia nested in Gaillardiinae, near Pelucha S. Watson, Psathyrotes, and Trichoptilium.

Selected References

None.

Lower Taxa

not tailed +, rounded +  and obtuse +
cream +  and purple +
connate +  and distinct +
subequal +
scarious +
usually ovate +  and lanceolate +
usually deltate +  and lanceolate +
corymbiform +  and open +
indeterminate +  and determinate +
Theodore M. Barkley† +, Luc Brouillet +  and John L. Strother +
H. Robinson +
decurrent +
3-nerved +  and 1-nerved +
spatulate;ovate;spatulate;ovate;oblanceolate;linear +
winged;ribbed;winged;ribbed +
tuberculate +  and rugose +
barbellulate +  and barbellate +
sessile +  and petiolate +
white +, purple +, pink +  and lavender +
5(-6)-angled +  and obpyramidal +
fertile +  and bisexual +
fertile +  and bisexual +
c +  and e United States +
pistillate +, staminate +  and neuter +
winged +  and beaked +
dispersed +
in open , corymbiform arrays +  and singly +
indeterminate +
homogamous +  and heterogamous +
each +  and sessile +
turbinate;obconic or hemispheric +
sessile +  and petiolate +
alternate +, whorled +  and opposite +
basal +  and cauline +
lance-linear +  and linear-oblong +
2-carpellate +
persistent +
10 +  and 20 +
styliferous +, neuter +  and pistillate +
bearing subulate enations +, hairy +  and bristly +
convex;conic +
lance-deltate +  and lance-attenuate +
exalbuminous +
prostrate +  and ascending +
appendaged +  and truncate +
hirsutulous +  and glabrous +
papillate +
Compositae +
Asteraceae (tribe Heliantheae) subtribe Marshalliinae +
Asteraceae tribe Heliantheae +
subtribe +
80 cm800 mm <br />0.8 m <br /> (120 cm1,200 mm <br />1.2 m <br />) +
20 cm200 mm <br />0.2 m <br /> (80 cm800 mm <br />0.8 m <br />) +
tree +, shrub +  and subshrub +