Athyrium filix-femina var. californicum

Butters

Rhodora 19: 201. 1918.

Common names: Southwestern lady fern
Endemic
Synonyms: Athyrium filix-femina subsp. californicum (Butters) Hultén
Treatment appears in FNA Volume 2.

Petiole 13–60 cm; scales brown or black, lanceolate or ovatelanceolate, 7–15 × 1–4 mm. Blade narrowly elliptic to oblanceolate, 1–2-pinnate-pinnatifid, 25–130 × 8–25 cm, gradually narrowed proximally, broadest at or just above middle, apex acuminate. Pinnae sessile or very short-stalked, linear-oblong, apex acuminate. Pinnules linear-oblong or linear-lanceolate, base basiscopically cuneate, acroscopically truncate, ± auriculate, apex ± acute to acuminate. Rachis, costae, and costules with scales and usually with pale glands. Sori elliptic or oblong, straight, hooked at distal end, or horseshoe-shaped, medial; indusia dentate or ciliate with nonglandular marginal hairs as long as width of indusia. Spores brown.


Habitat: Moist woods, meadows, streambanks
Elevation: 1000–3500 m

Distribution

V2 673-distribution-map.gif

Ariz., Calif., Colo., Idaho, Nev., N.Mex., Oreg., S.Dak., Utah, Wyo.

Discussion

This southwestern variety, Athyrium filix-femina var. californicum, occurs at higher elevations than var. cyclosorum. Athyrium filix-femina var. californicum and var. cyclosorum are more closely related to each other than to eastern varieties. Distinctness between western varieties was shown by F. K. Butters (1917); F. S. Liew (1972) treated them as consubspecific.

Selected References

None.

Lower Taxa

None.
Masahiro Kato +
Butters +
black;dark redbrown +
persisting +
auriculate +  and cuneate +
Polypodium filix-femina +
simple to commonly +
25 cm250 mm <br />0.25 m <br /> (?) +  and 130 cm1,300 mm <br />1.3 m <br /> (?) +
narrowed;1-2-pinnate-pinnatifid;narrowly elliptic;oblanceolate +
reduced +
broadest;8cm;25cm +
roundish +
Southwestern lady fern +
Ariz. +, Calif. +, Colo. +, Idaho +, Nev. +, N.Mex. +, Oreg. +, S.Dak. +, Utah +  and Wyo. +
1000–3500 m +
often +  and glabrous +
Moist woods, meadows, streambanks +
ciliate +  and dentate +
monomorphic +
straw-colored +
0.5 times length of blade;0.5 times length of blade +
13 cm130 mm <br />0.13 m <br /> (60 cm600 mm <br />0.6 m <br />) +
short-stalked +  and sessile +
linear-lanceolate +  and linear-oblong +
sessile +  and short-petiolulate +
with glands or hairs +  and glabrous +
round +, cuplike +, hoodlike +, reniform +, falcate +  and linear +
black;brown +
persistent +
0.7 cm7 mm <br />0.007 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
ovatelanceolate;lanceolate +
1mm +  and 4mm +
crenate +  and serrulate +
straight +
persistent +
with narrow sinus +  and attached +
discrete +
horseshoe--shaped +, hooked +, oblong +  and elliptic +
reniform +  and oblong +
dictyostelic +, unbranched +  and branched +
ascending +  and short-creeping +
Athyrium filix-femina subsp. californicum +
Athyrium filix-femina var. californicum +
Athyrium filix-femina +
variety +
epiphytic +  and hemiepiphytic +