Boechera breweri

(S. Watson) Al-Shehbaz

Novon 13: 384. 2003.

Basionym: Arabis breweri S. Watson Proc. Amer. Acad. Arts 11: 123. 1876
Synonyms: Arabis breweri var. figularis Jepson Arabis epilobioides
Treatment appears in FNA Volume 7. Treatment on page 367. Mentioned on page 354, 363, 365, 395, 404.

Perennials; long-lived; sexual; caudex woody. Stems usually 1 per caudex branch, arising from center of rosette, somewhat elevated on woody base or from ground surface, 0.6–3.5 (–4.5) dm, densely pubescent proximally, trichomes long-stalked and simple, 2-rayed (sometimes all simple), 0.4–1 mm, sparsely pubescent or glabrous distally. Basal leaves: blade oblanceolate, 3–7 (–11) mm wide, margins entire or dentate, ciliate proximally, trichomes to 1 mm, surfaces densely pubescent, trichomes long-stalked, 2–4-rayed, 0.4–0.8 mm. Cauline leaves: 5–15 (–28), often concealing stem proximally; blade auricles 0.5–3 (–5) mm, surfaces of distalmost leaves pubescent. Racemes 7–20 (–30) -flowered, usually unbranched. Fruiting pedicels ascending to divaricate-ascending, straight, 3–25 mm, pubescent or glabrous, trichomes spreading, simple and 2-rayed. Flowers ascending at anthesis; sepals pubescent; petals usually purple (rarely lavender), 7–12 × 2–4 mm, glabrous; pollen ellipsoid. Fruits ascending to divaricate-ascending, not appressed to rachis, not secund, usually curved, rarely straight, edges parallel, 3.5–10 cm × 1.5–2.2 mm; valves usually glabrous, rarely sparsely pubescent; ovules 48–96 per ovary; style 0.05–0.3 mm. Seeds uniseriate, 1.2–1.7 × 1–1.2 mm; wing continuous, 0.1–0.2 mm wide distally.

Discussion

Subspecies 2 (2 in the flora).

Relatively few other sexual diploid species of Boechera co-occur with B. breweri, and the only currently recognized hybrid involving this taxon is B. rubicundula (= B. arcuata × B. breweri; M. D. Windham and I. A. Al-Shehbaz 2007). Boechera breweri appears to be related to B. sparsiflora, and some specimens of subsp. shastaensis can be difficult to separate from that species. The two subspecies appear to intergrade on the western side of the Sacramento Valley from Tehama to Solano counties.

Selected References

None.

Key

1 Stems 0.6-2 dm, pubescent distally; fruiting pedicels 3-8(-12) mm, pubescent; fruits 3.5-7.5 cm. Boechera breweri subsp. breweri
1 Stems 1.8-3.5(-4.5) dm, glabrous distally; fruiting pedicels (7-)10-25 mm, usually glabrous; fruits (4.5-)7-10 cm. Boechera breweri subsp. shastaensis

"elongated" is not a number."thick" is not a number."dm" is not declared as a valid unit of measurement for this property.

... more about "Boechera breweri"
oblong +  and ovate +
Ihsan A. Al-Shehbaz +  and Michael D. Windham +
(S. Watson) Al-Shehbaz +
not +  and rosulate +
Arabis breweri +
appendaged +  and unappendaged +
0.3 cm3 mm <br />0.003 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
lyrate-pinnatifid +, dentate +  and entire +
dissected +, repand +, sinuate +, crenate +  and dentate +
oblanceolate +
much smaller +  and reduced +
3mm;7mm +
urceolate +, campanulate +  and tubular +
branched +  and simple +
petiolate +  and sessile +
well-developed +
indistinct +
not +  and differentiated +
emarginate +  and entire +
Calif. +  and Oreg. +
3.5 cm35 mm <br />0.035 m <br /> (10 cm100 mm <br />0.1 m <br />) +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.22 cm2.2 mm <br />0.0022 m <br />) +
straight +  and curved +
not winged +  and unappendaged +
connate +  and distinct +
actinomorphic +
straight +  and curved +
ascending +  and divaricate-ascending +
torulose +, falcate +, lanceolate +, oblong +  and linear +
annular +  and semiannular +
sessile +  and petiolate +
alternate +, not +  and rosulate +
cauline +  and basal +
connate +  and distinct +
decurrent +
dentate +  and entire +
pinnatifid +
prominent +  and obscure +
tenuinucellate +, crassinucellate +  and bitegmic +
campylotropous +  and anatropous +
Magenta (?) +, Red (?) +, Yellowish (?) +, Purple (?) +, Lavender (?) +, Pink (?) +  and White (?) +
0.7 cm7 mm <br />0.007 m <br /> (1.2 cm12 mm <br />0.012 m <br />) +
rudimentary +
0.2 cm2 mm <br />0.002 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
trinucleate +  and 3(-11)-colpate +
ellipsoid +
unbranched;7-20(-30)-flowered +
elongated +
reflexed +, ascending +  and erect +
slender +
mucilaginous +
margined +, not winged +  and winged +
biseriate +, sub-biseriate +  and uniseriate +
not +  and mucilaginous +
White (?) +, Black (?) +, Brown (?) +  and Yellow (?) +
0.12 cm1.2 mm <br />0.0012 m <br /> (0.17 cm1.7 mm <br />0.0017 m <br />) +
orbicular +, oblong +  and flattened +
0.1 cm1 mm <br />0.001 m <br /> (0.12 cm1.2 mm <br />0.0012 m <br />) +
persistent +  and caducous +
distinct +
spreading +, ascending +  and erect +
oblong +  and ovate +
reduced +
latiseptate +, schizocarpic +, samaroid +  and lomentaceous +
indehiscent +  and dehiscent +
angustiseptate +, flat +, angled +, terete +, torulose +, not +, segmented +  and nutletlike +
tetradynamous +
branched +  and unbranched +
procumbent +  and decumbent +
subsessile +, sessile +  and petiolate +
persistent +
distinct +
0.005 cm0.05 mm <br />5.0e-5 m <br /> (0.03 cm0.3 mm <br />3.0e-4 m <br />) +
Arabis breweri var. figularis +  and Arabis epilobioides +
Boechera breweri +
Boechera +
species +
simple +  and long-stalked +
glabrous +  and pubescent +
0.04 cm0.4 mm <br />4.0e-4 m <br /> (0.08 cm0.8 mm <br />8.0e-4 m <br />) +
coiled +, 1-7-veined +  and veined +
pubescent +  and glabrous +
anastomosing +
perennial +  and long-lived +
ascending +  and divaricate-ascending +
glabrous +  and pubescent +
0.3 cm3 mm <br />0.003 m <br /> (2.5 cm25 mm <br />0.025 m <br />) +
rhizomatous +, taprooted +, scapose +  and not +
perennial +, biennial +  and annual +
subshrub +  and herb +
aquatics +  and terrestrial +
glabrous +  and pubescent +
0.1mm;0.2mm +