Boechera californica

(Rollins) Windham & Al-Shehbaz

Harvard Pap. Bot. 11: 260. 2007.

Basionym: Arabis sparsiflora var. californica Rollins Rhodora 43: 402, fig. 14. 1941
Treatment appears in FNA Volume 7. Treatment on page 369. Mentioned on page 357, 358, 362, 407.

Perennials; usually long-lived; apomictic; caudex woody. Stems usually 1 per caudex branch, arising from center of rosette, elevated above ground surface on woody base, (2–) 3.5–13 dm, densely pubescent proximally, trichomes short-stalked, 2–4-rayed, 0.3–0.9 mm, moderately to sparsely pubescent distally. Basal leaves: blade linear to oblanceolate, 3–10 (–13) mm wide, margins entire or denticulate, ciliate along petiole, trichomes to 1.3 mm, surfaces densely pubescent, trichomes short-stalked, 4–8-rayed, 0.2–0.5 mm. Cauline leaves: 12–35 (–55), often concealing stem proximally; blade auricles 1–5 (–6) mm, surfaces of distalmost leaves pubescent. Racemes 30–120-flowered, usually unbranched. Fruiting pedicels descending to horizontal, usually recurved, rarely straight, 4–20 mm, pubescent, trichomes appressed, 3–7-rayed. Flowers divaricate-ascending at anthesis; sepals pubescent; petals usually purple, rarely pinkish, 9–14 × 1.5–3 mm, glabrous or sparsely pubescent (trichomes scattered abaxially); pollen spheroid. Fruits widely pendent, not appressed to rachis, not secund, usually curved, rarely straight, edges parallel, (6–) 8–12 cm × 1.5–2.5 mm; valves glabrous or trichomes sometimes scattered; ovules 140–180 per ovary; style 0.05–0.3 mm. Seeds uniseriate or sub-biseriate, 1.7–2 × 1.2–1.7 mm; wing continuous, 0.2–0.4 mm wide.


Phenology: Flowering Mar–Jun.
Habitat: Rocky slopes and gravelly soil in desert chaparral and oak woodlands
Elevation: 300-2300 m

Distribution

V7 501-distribution-map.gif

Calif., Mexico (Baja California)

Discussion

Morphological evidence suggests that Boechera californica is an apomictic species that arose through hybridization between B. arcuata and B. pulchra (see M. D. Windham and I. A. Al-Shehbaz 2007 for detailed comparison). In the flora area, B. californica is known only from Los Angeles, Riverside, San Bernardino, and San Diego counties.

Selected References

None.

Lower Taxa

None.

"elongated" is not a number."thick" is not a number."dm" is not declared as a valid unit of measurement for this property."dm" is not declared as a valid unit of measurement for this property.

... more about "Boechera californica"
oblong +  and ovate +
Ihsan A. Al-Shehbaz +  and Michael D. Windham +
(Rollins) Windham & Al-Shehbaz +
not +  and rosulate +
Arabis sparsiflora var. californica +
appendaged +  and unappendaged +
0.5 cm5 mm <br />0.005 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
0.1 cm1 mm <br />0.001 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
not +  and auriculate +
lyrate-pinnatifid +, dentate +  and entire +
dissected +, repand +, sinuate +, crenate +  and dentate +
linear;oblanceolate +
much smaller +  and reduced +
0.3 cm3 mm <br />0.003 m <br /> (1 cm10 mm <br />0.01 m <br />) +
urceolate +, campanulate +  and tubular +
branched +  and simple +
petiolate +  and sessile +
well-developed +
indistinct +
not +  and differentiated +
emarginate +  and entire +
Calif. +  and Mexico (Baja California) +
6 cm60 mm <br />0.06 m <br /> (8 cm80 mm <br />0.08 m <br />) +
8 cm80 mm <br />0.08 m <br /> (12 cm120 mm <br />0.12 m <br />) +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.25 cm2.5 mm <br />0.0025 m <br />) +
300-2300 m +
straight +  and curved +
not winged +  and unappendaged +
connate +  and distinct +
actinomorphic +
divaricate-ascending +
straight +  and curved +
torulose +, falcate +, lanceolate +, oblong +  and linear +
Rocky slopes and gravelly soil in desert chaparral and oak woodlands +
annular +  and semiannular +
sessile +  and petiolate +
alternate +, not +  and rosulate +
cauline +  and basal +
connate +  and distinct +
decurrent +
dentate +  and entire +
denticulate;entire +
prominent +  and obscure +
tenuinucellate +, crassinucellate +  and bitegmic +
campylotropous +  and anatropous +
9mm;14mm +
rudimentary +
pubescent +  and glabrous +
oblanceolate +  and spatulate +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
Flowering Mar–Jun. +
trinucleate +  and 3(-11)-colpate +
spheroid +
Harvard Pap. Bot. +
unbranched;30-120-flowered +
reflexed +, ascending +  and erect +
slender +
mucilaginous +
margined +, not winged +  and winged +
sub-biseriate +  and uniseriate +
not +  and mucilaginous +
white +, black +, brown +  and yellow +
0.17 cm1.7 mm <br />0.0017 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
orbicular +, oblong +  and flattened +
0.12 cm1.2 mm <br />0.0012 m <br /> (0.17 cm1.7 mm <br />0.0017 m <br />) +
persistent +  and caducous +
distinct +
spreading +, ascending +  and erect +
oblong +  and ovate +
reduced +
latiseptate +, schizocarpic +, samaroid +  and lomentaceous +
indehiscent +  and dehiscent +
angustiseptate +, flat +, angled +, terete +, torulose +, not +, segmented +  and nutletlike +
tetradynamous +
branched +  and unbranched +
procumbent +  and decumbent +
subsessile +, sessile +  and petiolate +
persistent +
distinct +
0.005 cm0.05 mm <br />5.0e-5 m <br /> (0.03 cm0.3 mm <br />3.0e-4 m <br />) +
Cruciferae +
Boechera californica +
Boechera +
species +
0.02 cm0.2 mm <br />2.0e-4 m <br /> (0.05 cm0.5 mm <br />5.0e-4 m <br />) +
coiled +, 1-7-veined +  and veined +
anastomosing +
perennial +  and long-lived +
recurved +, descending +  and horizontal +
0.4 cm4 mm <br />0.004 m <br /> (2 cm20 mm <br />0.02 m <br />) +
rhizomatous +, taprooted +, scapose +  and not +
perennial +, biennial +  and annual +
subshrub +  and herb +
aquatics +  and terrestrial +
glabrous +  and pubescent +
0.2mm;0.4mm +