Boechera grahamii

(Lehmann) Windham & Al-Shehbaz

Harvard Pap. Bot. 12: 241. 2007.

Basionym: Turritis grahamii Lehmann Index Seminum (Hamburg) 1831: 7. 1831,
Synonyms: Arabis boivinii G. A. Mulligan Arabis bourgovii Rydberg Arabis confinis S. Watson Arabis confinis var. brachycarpa (Torrey & A. Gray) S. Watson & J. M. Coulter Arabis dacotica A. Nelson Arabis divaricarpa var. dacotica (Greene) B. Boivin Arabis divaricarpa var. hemicylindrica B. Boivin Arabis divaricarpa var. stenocarpa M. Hopkins Arabis drummondii var. brachycarpa (Torrey & A. Gray) A. Gray Arabis holboellii var. brachycarpa (Torrey & A. Gray) S. L. Welsh Boechera brachycarpa Turritis brachycarpa
Treatment appears in FNA Volume 7. Treatment on page 382. Mentioned on page 356, 359, 363, 374, 401.

Biennials or perennials; short-lived; apomictic; caudex present or absent. Stems usually 1 per caudex branch, arising from center of rosette near ground surface, (1.2–) 2–12 dm, glabrous or pubescent proximally, trichomes simple and sessile or subsessile, 2 or 3 (–6) -rayed, to 1 mm, glabrous distally. Basal leaves: blade oblanceolate, 1.5–10 (–20) mm wide, margins entire or dentate, ciliate proximally, trichomes (simple), to 1 mm, surfaces sparsely to densely pubescent, trichomes sessile or subsessile, 2–4 (–7) -rayed, 0.2–0.6 mm. Cauline leaves: (10–) 13–52, often concealing stem proximally; blade auricles 1–5 mm, surfaces of distalmost leaves glabrous. Racemes 16–88-flowered, usually unbranched. Fruiting pedicels divaricate-ascending to descending, usually gently curved downward, rarely straight, 5–22 (–30) mm, glabrous. Flowers divaricate at anthesis; sepals glabrous or pubescent; petals usually white, rarely lavender, 5.5–8 × 1–2 mm, glabrous; pollen spheroid. Fruits divaricate-ascending to pendent, not appressed to rachis, not secund, straight or slightly curved, edges parallel, 3.5–9 cm × 1–1.8 mm; valves glabrous; ovules 84–146 per ovary; style 0.05–0.5 mm. Seeds uniseriate to sub-biseriate, 1–1.8 × 0.8–1.2 mm; wing continuous, 0.1–0.25 mm wide. 2n = 21.


Phenology: Flowering May–Aug.
Habitat: Rocky slopes and sandy soil in prairies and open forests
Elevation: 100-3200 m

Distribution

V7 534-distribution-map.gif

Alta., B.C., Man., N.W.T., Ont., Que., Sask., Yukon, Alaska, Colo., Mich., Minn., Mont., N.Y., N.Dak., S.Dak., Vt., Wash., Wis., Wyo.

Discussion

Morphological evidence suggests that Boechera grahamii is an apomictic species that arose through hybridization between B. collinsii and B. stricta. Previous authors have assigned these specimens to Arabis (Boechera) divaricarpa (see M. D. Windham and I. A. Al-Shehbaz 2007b for detailed comparison); if these taxa are treated as conspecific, the name B. grahamii has priority.

Selected References

None.

Lower Taxa

None.

"elongated" is not a number."thick" is not a number."dm" is not declared as a valid unit of measurement for this property."dm" is not declared as a valid unit of measurement for this property.

... more about "Boechera grahamii"
oblong +  and ovate +
Ihsan A. Al-Shehbaz +  and Michael D. Windham +
(Lehmann) Windham & Al-Shehbaz +
not +  and rosulate +
Turritis grahamii +
appendaged +  and unappendaged +
0.1 cm1 mm <br />0.001 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
not +  and auriculate +
lyrate-pinnatifid +, dentate +  and entire +
dissected +, repand +, sinuate +, crenate +  and dentate +
oblanceolate +
much smaller +  and reduced +
0.15 cm1.5 mm <br />0.0015 m <br /> (1 cm10 mm <br />0.01 m <br />) +
urceolate +, campanulate +  and tubular +
branched +  and simple +
petiolate +  and sessile +
well-developed +
indistinct +
not +  and differentiated +
emarginate +  and entire +
Alta. +, B.C. +, Man. +, N.W.T. +, Ont. +, Que. +, Sask. +, Yukon +, Alaska +, Colo. +, Mich. +, Minn. +, Mont. +, N.Y. +, N.Dak. +, S.Dak. +, Vt. +, Wash. +, Wis. +  and Wyo. +
3.5 cm35 mm <br />0.035 m <br /> (9 cm90 mm <br />0.09 m <br />) +
0.1 cm1 mm <br />0.001 m <br /> (0.18 cm1.8 mm <br />0.0018 m <br />) +
100-3200 m +
straight +  and curved +
not winged +  and unappendaged +
connate +  and distinct +
actinomorphic +
curved +  and straight +
divaricate-ascending +  and pendent +
torulose +, falcate +, lanceolate +, oblong +  and linear +
Rocky slopes and sandy soil in prairies and open forests +
annular +  and semiannular +
sessile +  and petiolate +
alternate +, not +  and rosulate +
cauline +  and basal +
connate +  and distinct +
decurrent +
dentate +  and entire +
pinnatifid +
prominent +  and obscure +
tenuinucellate +, crassinucellate +  and bitegmic +
campylotropous +  and anatropous +
5.5mm;8mm +
rudimentary +
oblanceolate +  and spatulate +
0.1 cm1 mm <br />0.001 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
Flowering May–Aug. +
trinucleate +  and 3(-11)-colpate +
spheroid +
Harvard Pap. Bot. +
unbranched;16-88-flowered +
2.2 cm22 mm <br />0.022 m <br /> (3 cm30 mm <br />0.03 m <br />) +
straight +  and curved +
downward +, divaricate-ascending +  and descending +
slender +
0.5 cm5 mm <br />0.005 m <br /> (2.2 cm22 mm <br />0.022 m <br />) +
mucilaginous +
margined +, not winged +  and winged +
uniseriate +  and sub-biseriate +
not +  and mucilaginous +
white +, black +, brown +  and yellow +
0.1 cm1 mm <br />0.001 m <br /> (0.18 cm1.8 mm <br />0.0018 m <br />) +
orbicular +, oblong +  and flattened +
0.08 cm0.8 mm <br />8.0e-4 m <br /> (0.12 cm1.2 mm <br />0.0012 m <br />) +
persistent +  and caducous +
distinct +
spreading +, ascending +  and erect +
pubescent +  and glabrous +
oblong +  and ovate +
reduced +
latiseptate +, schizocarpic +, samaroid +  and lomentaceous +
indehiscent +  and dehiscent +
angustiseptate +, flat +, angled +, terete +, torulose +, not +, segmented +  and nutletlike +
tetradynamous +
branched +  and unbranched +
procumbent +  and decumbent +
pubescent +  and glabrous +
subsessile +, sessile +  and petiolate +
persistent +
distinct +
0.005 cm0.05 mm <br />5.0e-5 m <br /> (0.05 cm0.5 mm <br />5.0e-4 m <br />) +
glabrous +  and pubescent +
Arabis boivinii +, Arabis bourgovii +, Arabis confinis +, Arabis confinis var. brachycarpa +, Arabis dacotica +, Arabis divaricarpa var. dacotica +, Arabis divaricarpa var. hemicylindrica +, Arabis divaricarpa var. stenocarpa +, Arabis drummondii var. brachycarpa +, Arabis holboellii var. brachycarpa +, Boechera brachycarpa +  and Turritis brachycarpa +
Boechera grahamii +
Boechera +
species +
subsessile +, sessile +  and simple +
3(-6)-rayed +  and 2 +
0.02 cm0.2 mm <br />2.0e-4 m <br /> (0.06 cm0.6 mm <br />6.0e-4 m <br />) +
coiled +, 1-7-veined +  and veined +
anastomosing +
perennial +, short-lived +  and biennial +
hispid +, hirsute +, pubescent +  and glabrous +
rhizomatous +, taprooted +, scapose +  and not +
perennial +, biennial +  and annual +
subshrub +  and herb +
aquatics +  and terrestrial +
glabrous +  and pubescent +
0.1mm;0.25mm +