Boechera puberula

(Nuttall) Dorn

Brittonia, 55: 3. 2003.

Basionym: Arabis puberula Nuttall in J. Torrey and A. Gray, Fl. N. Amer. 1: 82. 1838
Synonyms: Arabis arida Greene Arabis beckwithii S. Watson Arabis lignipes var. impar A. Nelson Arabis sabulosa M. E. Jones Arabis subpinnatifida var. beckwithii (S. Watson) Jepson Arabis subpinnatifida var. impar (A. Nelson) Rollins Boechera beckwithii
Treatment appears in FNA Volume 7. Treatment on page 399. Mentioned on page 351, 354, 386, 400, 409.

Perennials; short-lived; sexual; caudex not woody. Stems usually 1 per caudex branch, arising from center of rosette near ground surface, (1–) 2–6.3 dm, densely pubescent proximally, trichomes short-stalked, 3–8-rayed, 0.1–0.3 (–0.5) mm, similarly pubescent distally. Basal leaves: blade oblanceolate to linear-oblanceolate, 1.5–5 mm wide, margins usually dentate, not ciliate, surfaces densely pubescent, trichomes short-stalked, 5–12-rayed, 0.05–0.2 mm. Cauline leaves: 7–45 (–65), concealing stem proximally; blade auricles absent or 0.7–3 mm, surfaces of distalmost leaves pubescent. Racemes 10–40 (–64) -flowered, usually unbranched. Fruiting pedicels pendent, recurved distal to horizontal to ascending base, 4–10 mm, pubescent, trichomes appressed, branched. Flowers ascending to descending at anthesis; sepals pubescent; petals white to lavender, 5–9 × 0.8–1.8 mm, glabrous; pollen ellipsoid. Fruits closely pendent, rarely appressed to rachis, sometimes somewhat secund, usually straight, edges parallel, 3–6.5 cm × 1.9–2.2 mm; valves pubescent throughout; ovules 38–64 per ovary; style 0.05–0.1 mm. Seeds uniseriate, 1.4–1.8 × 1–1.4 mm; wing continuous, 0.1–0.3 mm wide. 2n = 14.


Phenology: Flowering Apr–Jul.
Habitat: Ledges, rocky slopes, gravelly hillsides in sagebrush, pinyon-juniper, and mountain shrub communities
Elevation: 1300-2900 m

Distribution

V7 574-distribution-map.gif

Calif., Nev., Oreg., Utah.

Discussion

Boechera puberula is a diploid species that appears to intergrade with both B. retrofracta and B. subpinnatifida. The glabrous-fruited specimens discussed by R. C. Rollins (1993) represent apomictic hybrids with other species, primarily B. pendulocarpa.

Selected References

None.

Lower Taxa

None.

"elongated" is not a number."thick" is not a number."dm" is not declared as a valid unit of measurement for this property."dm" is not declared as a valid unit of measurement for this property.

... more about "Boechera puberula"
oblong +  and ovate +
Ihsan A. Al-Shehbaz +  and Michael D. Windham +
(Nuttall) Dorn +
not +  and rosulate +
horizontal +  and ascending +
0.4 cm4 mm <br />0.004 m <br /> (1 cm10 mm <br />0.01 m <br />) +
Arabis puberula +
appendaged +  and unappendaged +
not +  and auriculate +
lyrate-pinnatifid +, dentate +  and entire +
dissected +, repand +, sinuate +, crenate +  and dentate +
oblanceolate;linear-oblanceolate +
much smaller +  and reduced +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
urceolate +, campanulate +  and tubular +
branched +  and simple +
not woody +
petiolate +  and sessile +
well-developed +
indistinct +
not +  and differentiated +
emarginate +  and entire +
Calif. +, Nev. +, Oreg. +  and Utah. +
3 cm30 mm <br />0.03 m <br /> (6.5 cm65 mm <br />0.065 m <br />) +
0.19 cm1.9 mm <br />0.0019 m <br /> (0.22 cm2.2 mm <br />0.0022 m <br />) +
1300-2900 m +
straight +  and curved +
not winged +  and unappendaged +
connate +  and distinct +
actinomorphic +
ascending +  and descending +
straight +
torulose +, falcate +, lanceolate +, oblong +  and linear +
Ledges, rocky slopes, gravelly hillsides in sagebrush, pinyon-juniper, and mountain shrub communities +
annular +  and semiannular +
sessile +  and petiolate +
alternate +, not +  and rosulate +
cauline +  and basal +
connate +  and distinct +
decurrent +
pinnatifid +
prominent +  and obscure +
tenuinucellate +, crassinucellate +  and bitegmic +
campylotropous +  and anatropous +
white +  and lavender +
rudimentary +
oblanceolate +  and spatulate +
0.08 cm0.8 mm <br />8.0e-4 m <br /> (0.18 cm1.8 mm <br />0.0018 m <br />) +
Flowering Apr–Jul. +
trinucleate +  and 3(-11)-colpate +
ellipsoid +
unbranched;10-40(-64)-flowered +
reflexed +, ascending +  and erect +
slender +
mucilaginous +
margined +, not winged +  and winged +
biseriate +, sub-biseriate +  and uniseriate +
not +  and mucilaginous +
white +, black +, brown +  and yellow +
0.14 cm1.4 mm <br />0.0014 m <br /> (0.18 cm1.8 mm <br />0.0018 m <br />) +
orbicular +, oblong +  and flattened +
0.1 cm1 mm <br />0.001 m <br /> (0.14 cm1.4 mm <br />0.0014 m <br />) +
persistent +  and caducous +
distinct +
spreading +, ascending +  and erect +
oblong +  and ovate +
reduced +
latiseptate +, schizocarpic +, samaroid +  and lomentaceous +
indehiscent +  and dehiscent +
angustiseptate +, flat +, angled +, terete +, torulose +, not +, segmented +  and nutletlike +
tetradynamous +
branched +  and unbranched +
procumbent +  and decumbent +
subsessile +, sessile +  and petiolate +
persistent +
distinct +
0.005 cm0.05 mm <br />5.0e-5 m <br /> (0.01 cm0.1 mm <br />1.0e-4 m <br />) +
Arabis arida +, Arabis beckwithii +, Arabis lignipes var. impar +, Arabis sabulosa +, Arabis subpinnatifida var. beckwithii +, Arabis subpinnatifida var. impar +  and Boechera beckwithii +
Boechera puberula +
Boechera +
species +
branched +  and short-stalked +
0.03 cm0.3 mm <br />3.0e-4 m <br /> (0.05 cm0.5 mm <br />5.0e-4 m <br />) +
0.005 cm0.05 mm <br />5.0e-5 m <br /> (0.02 cm0.2 mm <br />2.0e-4 m <br />) +
coiled +, 1-7-veined +  and veined +
anastomosing +
perennial +  and short-lived +
hispid +, hirsute +, pubescent +  and glabrous +
rhizomatous +, taprooted +, scapose +  and not +
perennial +, biennial +  and annual +
subshrub +  and herb +
aquatics +  and terrestrial +
glabrous +  and pubescent +
0.1mm;0.3mm +