Boechera subpinnatifida

(S. Watson) Al-Shehbaz

Novon 13: 389. 2003.

Basionym: Arabis subpinnatifida S. Watson Proc. Amer. Acad. Arts 20: 353. 1885
Treatment appears in FNA Volume 7. Treatment on page 409. Mentioned on page 354, 357, 400, 406.

Perennials; long-lived; sexual; caudex woody (often with persistent, crowded leaf-bases). Stems usually 1 per caudex branch, arising from center of rosette near ground surface, 1–4 (–5) dm, densely pubescent proximally, trichomes short-stalked, 2–6-rayed, 0.1–0.2 mm, sparsely pubescent distally. Basal leaves: blade narrowly oblanceolate, 1–4 (–5) mm wide, margins prominently dentate to subpinnatifid (leaf margins of sterile shoots often entire), ciliate near petiole base, trichomes (simple or 2-rayed), 0.4–0.6 mm, surfaces densely pubescent, trichomes short-stalked, (2–) 4–9-rayed, 0.05–0.2 mm. Cauline leaves: (10–) 20–60, often concealing stem throughout; blade auricles 0.5–3 mm, surfaces of distalmost leaves moderately to sparsely pubescent. Racemes 8–30-flowered, usually unbranched. Fruiting pedicels reflexed, strongly recurved, 5–15 mm, pubescent, trichomes appressed, branched. Flowers divaricate-ascending to pendent at anthesis; sepals pubescent; petals usually purple, rarely lavender, 9–14 × 1.5–3 mm, glabrous; pollen ellipsoid. Fruits pendent, not appressed to rachis, not secund, straight to slightly curved, edges parallel, (3.5–) 5–8 cm × (1.6–) 2–3 mm; valves pubescent throughout; ovules 24–42 per ovary; style 0.5–1 mm. Seeds uniseriate, 2.5–3.5 × 1.5–2.2 mm; wing continuous or at both ends, 0.4–0.8 mm wide. 2n = 14.


Phenology: Flowering Mar–May.
Habitat: Rock outcrops, talus, gravelly soil, often in sagebrush-grassland communities
Elevation: 800-2400 m

Distribution

V7 595-distribution-map.gif

Calif., Idaho, Nev., Oreg., Utah.

Discussion

Originally thought to be restricted to northern California and adjacent Oregon, Boechera subpinnatifida is a sexual species that recently has been found in central Idaho, northern Nevada, and northwestern Utah. It appears to intergrade with both B. puberula and B. retrofracta, and species boundaries within this complex need further study.

Selected References

None.

Lower Taxa

None.

"elongated" is not a number."thick" is not a number."dm" is not declared as a valid unit of measurement for this property.

... more about "Boechera subpinnatifida"
oblong +  and ovate +
Ihsan A. Al-Shehbaz +  and Michael D. Windham +
(S. Watson) Al-Shehbaz +
not +  and rosulate +
Arabis subpinnatifida +
appendaged +  and unappendaged +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
lyrate-pinnatifid +, dentate +  and entire +
dissected +, repand +, sinuate +, crenate +  and dentate +
oblanceolate +
much smaller +  and reduced +
1mm;4mm +
urceolate +, campanulate +  and tubular +
branched +  and simple +
petiolate +  and sessile +
well-developed +
indistinct +
not +  and differentiated +
emarginate +  and entire +
Calif. +, Idaho +, Nev. +, Oreg. +  and Utah. +
3.5 cm35 mm <br />0.035 m <br /> (5 cm50 mm <br />0.05 m <br />) +
0.16 cm1.6 mm <br />0.0016 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
5 cm50 mm <br />0.05 m <br /> (8 cm80 mm <br />0.08 m <br />) +
0.2 cm2 mm <br />0.002 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
800-2400 m +
straight +  and curved +
not winged +  and unappendaged +
connate +  and distinct +
actinomorphic +
divaricate-ascending +  and pendent +
straight +  and slightly curved +
torulose +, falcate +, lanceolate +, oblong +  and linear +
Rock outcrops, talus, gravelly soil, often in sagebrush-grassland communities +
annular +  and semiannular +
sessile +  and petiolate +
alternate +, not +  and rosulate +
cauline +  and basal +
connate +  and distinct +
decurrent +
prominently dentate;subpinnatifid +
dentate +  and entire +
pinnatifid +
prominent +  and obscure +
tenuinucellate +, crassinucellate +  and bitegmic +
campylotropous +  and anatropous +
Magenta (?) +, Red (?) +, Yellowish (?) +, Purple (?) +, Lavender (?) +, Pink (?) +  and White (?) +
0.9 cm9 mm <br />0.009 m <br /> (1.4 cm14 mm <br />0.014 m <br />) +
rudimentary +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
Flowering Mar–May. +
trinucleate +  and 3(-11)-colpate +
ellipsoid +
unbranched;8-30-flowered +
elongated +
reflexed +, ascending +  and erect +
slender +
mucilaginous +
margined +, not winged +  and winged +
biseriate +, sub-biseriate +  and uniseriate +
not +  and mucilaginous +
White (?) +, Black (?) +, Brown (?) +  and Yellow (?) +
0.25 cm2.5 mm <br />0.0025 m <br /> (0.35 cm3.5 mm <br />0.0035 m <br />) +
orbicular +, oblong +  and flattened +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.22 cm2.2 mm <br />0.0022 m <br />) +
persistent +  and caducous +
distinct +
spreading +, ascending +  and erect +
oblong +  and ovate +
reduced +
latiseptate +, schizocarpic +, samaroid +  and lomentaceous +
indehiscent +  and dehiscent +
angustiseptate +, flat +, angled +, terete +, torulose +, not +, segmented +  and nutletlike +
tetradynamous +
branched +  and unbranched +
procumbent +  and decumbent +
subsessile +, sessile +  and petiolate +
persistent +
distinct +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
Cruciferae +
Boechera subpinnatifida +
Boechera +
species +
branched +  and short-stalked +
0.005 cm0.05 mm <br />5.0e-5 m <br /> (0.02 cm0.2 mm <br />2.0e-4 m <br />) +
coiled +, 1-7-veined +  and veined +
anastomosing +
perennial +  and long-lived +
recurved +  and reflexed +
0.5 cm5 mm <br />0.005 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
rhizomatous +, taprooted +, scapose +  and not +
perennial +, biennial +  and annual +
subshrub +  and herb +
aquatics +  and terrestrial +
glabrous +  and pubescent +
at both ends +  and continuous +
0.4mm;0.8mm +