Boltonia asteroides var. asteroides

IllustratedEndemic
Synonyms: Boltonia asteroides var. glastifolia Boltonia glastifolia
Treatment appears in FNA Volume 20. Treatment on page 355. Mentioned on page 356.

Heads 10–100+; bracts linear-oblanceolate to narrowly oblanceolate, (4–) 15–90 × 2–9 mm. Involucres 2.4–3.8 × 3.7–8.7 mm. Phyllaries linear to linear-attenuate or subulate, subequal, membranous margins 0.5–1 mm wide, apices acute; outer 1.7–2.8 × 0.4–0.6 (–0.9) mm; inner 2.2–3.9 × 0.3–0.7 (–1) mm. Cypselae obovoid, sometimes wingless; pappus awns (0–) 0.4–1.2 mm, to 2/3 length of cypselae. 2n = 36.


Phenology: Flowering Jul–Nov.
Habitat: Marshes, wet prairies, roadsides, wet pine savannas, riverbanks, flood plains, swales
Elevation: 0–400 m

Distribution

V20-820-distribution-map.gif

Ala., Ark., Del., Fla., Ga., La., Md., Miss., N.J., N.Y., N.C., Pa., S.C., Tex., Va.

Discussion

Variety asteroides is considered endangered in Maryland and Pennsylvania. Populations of var. asteroides restricted to the sinkholes of Virginia and New Jersey are atypical in cypsela morphology. They lack pappus awns and wings are reduced or almost absent, characters typical of the cypselae of Boltonia caroliniana. Other characters (arrays with stiff, ascending branches, linear to linear-attenuate, subequal phyllaries, size of leaves and heads) are characteristic of var. asteroides elsewhere. Possibly ancient hybridization between B. caroliniana and var. asteroides may have occurred. Further detailed studies of the populations may help in resolving their taxonomic status.

Selected References

None.

Lower Taxa

None.

not tailed +, rounded +  and obtuse +
distinct +
subequal +
scarious +
usually triangular +  and linear +
usually deltate +  and lanceolate +
paniculiform +  and corymbiform +
indeterminate +  and determinate +
Vesna Karaman-Castro +  and Lowell E. Urbatsch +
(Linnaeus) L’Héritier +
7 +  and 12 +
not decurrent +
Matricaria asteroides +
subulate;lanceolate or oblanceolate +
rugulose +  and muricate +
0.4 cm4 mm <br />0.004 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
1.2 cm12 mm <br />0.012 m <br /> (6.2 cm62 mm <br />0.062 m <br />) +
1.5 cm15 mm <br />0.015 m <br /> (9 cm90 mm <br />0.09 m <br />) +
linear-oblanceolate +  and narrowly oblanceolate +
0.2 cm2 mm <br />0.002 m <br /> (0.9 cm9 mm <br />0.009 m <br />) +
spreading;ascending +
barbellulate +  and barbellate +
2 cm20 mm <br />0.02 m <br /> (22 cm220 mm <br />0.22 m <br />) +
0.2 cm2 mm <br />0.002 m <br /> (3 cm30 mm <br />0.03 m <br />) +
not 2-lipped +  and actinomorphic +
white +  and lilac +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.26 cm2.6 mm <br />0.0026 m <br />) +
beaked +, 2-ribbed +  and 5-ribbed +
0.1 cm1 mm <br />0.001 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
0.08 cm0.8 mm <br />8.0e-4 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
fertile +  and bisexual +
fertile +  and bisexual +
compressed;obovoid;cuneate or obcordiform +
Ala. +, Ark. +, Del. +, Fla. +, Ga. +, La. +, Md. +, Miss. +, N.J. +, N.Y. +, N.C. +, Pa. +, S.C. +, Tex. +  and Va. +
0–400 m +
puberulent +  and glabrous +
pistillate +, staminate +  and neuter +
winged +  and beaked +
dispersed +
Marshes, wet prairies, roadsides, wet pine savannas, riverbanks, flood plains, swales +
indeterminate +
heterogamous +  and homogamous +
each +  and sessile +
0.07 cm0.7 mm <br />7.0e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
0.22 cm2.2 mm <br />0.0022 m <br /> (0.39 cm3.9 mm <br />0.0039 m <br />) +
0.03 cm0.3 mm <br />3.0e-4 m <br /> (0.07 cm0.7 mm <br />7.0e-4 m <br />) +
0.24 cm2.4 mm <br />0.0024 m <br /> (0.38 cm3.8 mm <br />0.0038 m <br />) +
hemispheric +
0.37 cm3.7 mm <br />0.0037 m <br /> (0.87 cm8.7 mm <br />0.0087 m <br />) +
0.5 cm5 mm <br />0.005 m <br /> (1.3 cm13 mm <br />0.013 m <br />) +
cauline +  and basal +
triangular +
not +  and winged +
entire +  and serrulate +
0.5mm +  and 1mm +
0.06 cm0.6 mm <br />6.0e-4 m <br /> (0.09 cm0.9 mm <br />9.0e-4 m <br />) +
0.17 cm1.7 mm <br />0.0017 m <br /> (0.28 cm2.8 mm <br />0.0028 m <br />) +
0.04 cm0.4 mm <br />4.0e-4 m <br /> (0.06 cm0.6 mm <br />6.0e-4 m <br />) +
2-carpellate +
0.04 cm0.4 mm <br />4.0e-4 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
0 cm0 mm <br />0 m <br /> (0.04 cm0.4 mm <br />4.0e-4 m <br />) +
0.04 cm0.4 mm <br />4.0e-4 m <br /> (0.12 cm1.2 mm <br />0.0012 m <br />) +
persistent +
0.5 cm5 mm <br />0.005 m <br /> (22 cm220 mm <br />0.22 m <br />) +
Flowering Jul–Nov. +
linear +  and linear-attenuate or subulate +
Sert. Angl., +
triquetrous +
bearing subulate enations +, hairy +  and bristly +
conic;hemispheric +
adaxial +  and abaxial +
1 +  and 2 +
exalbuminous +
Illustrated +  and Endemic +
appendaged +  and truncate +
papillate +  and smooth +
Boltonia asteroides var. glastifolia +  and Boltonia glastifolia +
Boltonia asteroides var. asteroides +
Boltonia asteroides +
variety +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.17 cm1.7 mm <br />0.0017 m <br />) +
30 cm300 mm <br />0.3 m <br /> (200 cm2,000 mm <br />2 m <br />) +
0.1mm +  and 0.5mm +