familyOphioglossaceae
genusBotrychium
subgenusBotrychium subg. Botrychium
speciesBotrychium paradoxum

Botrychium paradoxum

W. H. Wagner

Amer. Fern J. 71: 24. 1981.

Warren H. Wagner Jr., Florence S. Wagner
Common names: Paradox moonwort
IllustratedEndemicConservation concern
Treatment appears in FNA Volume 2.
Please click on the illustration for a higher resolution version.
Illustrator: John Myers

Copyright: Flora of North America Association

Trophophores converted entirely to second fertile segment, stalk 1/2 length of fertile segment. Sporophores double, 2 per leaf, 1-pinnate, 0.5–4 cm. 2n =180.


Habitat: Sporophores in June to August. Difficult to detect, plants usually hidden under other vegetation, in snowfields, secondary growth pastures
Elevation: 1500–3000 m

Distribution

V2 685-distribution-map.gif

Alta., B.C., Sask., Mont., Utah.

Discussion

The leaf structure of Botrychium paradoxum is uniform and unique. Very rare teratological individuals of other moonwort species may have trophophores partially or wholly transformed into sporophores.

Botrychium × watertonense W.H. Wagner, known only from one locality in western Alberta, is the sterile hybrid of B. hesperium and B. paradoxum. It can be identified by its trophophore pinnae; all are bordered with sporangia. It may reproduce by some unknown mechanism, such as unreduced spores (W.H. Wagner Jr., F. S. Wagner, et al. 1984).

Of conservation concern.

Selected References

None.

Lower Taxa

None.
... more about "Botrychium paradoxum"
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2n chromosome quantity
180 +
Apex shape
acute +  and rounded +
Author
Warren H. Wagner Jr. +  and Florence S. Wagner +
Authority
W. H. Wagner +
Blade architecture
simple +  and 5-pinnate +
Blade condition
dying +
Blade length
4 cm40 mm <br />0.04 m <br /> (25 cm250 mm <br />0.25 m <br />) +
Blade presence
absent +
Blade quantity
1 +
Blade shape
mostly linear +  and oblong +
Blade texture
herbaceous +  and leathery +
Blade width
0 cm0 mm <br />0 m <br /> (2.5 cm25 mm <br />0.025 m <br />) +
Common name
Paradox moonwort +
Distribution
Alta. +, B.C. +, Sask. +, Mont. +  and Utah. +
Elevation
1500–3000 m +
Gametophyte architecture
unbranched +
Gametophyte coloration
not green +
Gametophyte length
0.1 cm1 mm <br />0.001 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
Gametophyte location
subterranean +
Gametophyte shape
ovate +
Gametophyte structure in adjective form
mycorrhizal +
Gametophyte texture
fleshy +
Gametophyte width
0.1 cm1 mm <br />0.001 m <br /> (1 cm10 mm <br />0.01 m <br />) +
Gemma presence
absent +
Gemma shape
Spheric (?) +
Gemma size
minute +
Habitat
Sporophores in June to August. Difficult to detect, plants usually hidden under other vegetation, in snowfields, secondary growth pastures +
Hair architecture or arrangement
uniseriate +
Hair arrangement
scattered +
Hair length or size
long +
Idioblast quantity
lacking +
Illustration
Present +
Illustration copyright
Flora of North America Association +
Illustrator
John Myers +
Indument presence
absent +
Indument quantity
of widely scattered , long , uniseriate hairs +
Lateral branch quantity
few +
Lateral branch size or width
narrow +
Leaf-base architecture
open +
Leaf-base architecture or fixation
clasping +
Leaf-base fusion
fused +
Leaf-base shape
dilated +
Leaf atypical quantity
1 (2) +
Leaf primordium development
successive +
Leaf primordium position relational
surrounding +
Leaf primordium pubescence
glabrous +
Leaf quantity
1 (?) +
Leaf reproduction
fertile +
Margin architecture or shape
entire +
Margin shape
dentate;lacerate +
Number of lower taxa
0 +
Pinna lobe architecture or shape
asymmetric +
Pinna orientation
spreading;ascending +
Pinna segment architecture or shape
asymmetric +
Pinna shape
Fan-shaped (?) +  and Lanceolate (?) +
Plants perennial habitat
epiphytic +  and terrestrial +
Portion architecture
spore-bearing +  and laminate +
Portion nutrition
photosynthetic +
Portion reproduction
sterile +
Primordium pubescence
with long , uniseriate hairs +  and glabrous +
Publication title
Amer. Fern J. +
Publication year
1981 +
Rachis shape
flattened +
Rachis texture
not fleshy +
Rachis width
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
Reference
None +
Ridge pubescence or texture
corky +
Root architecture
branching +
Root architecture or pubescence or relief
with corky ridges +  and smooth +
Root coloration
list +  and count +
Root diameter
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
Root hair quantity
lacking +
Root location
1 +
Root quantity
10 +
Root reproduction
not proliferous +
Row quantity
2 +
Segment reproduction
fertile +
Sheath condition
closed +
Source xml
https://bibilujan@bitbucket.org/aafc-mbb/fna-fine-grained-xml.git/src/84153f6d59a0a91d69695978a64cee7560374f8e/V2/V2 685.xml +
Special status
Illustrated +, Endemic +  and Conservation concern +
Species architecture
branched +
Species quantity
1 +
Sporangium architecture
sessile +  and short-stalked +
Sporangium arrangement
Cluster (?) +
Sporangium diameter
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
Sporangium position relational
embedded +
Sporangium prominence
exposed +
Spore architecture
thick-walled +  and trilete +
Spore quantity
1 +
Sporophore architecture
short-stalked +
Sporophore architecture or shape
1-pinnate +
Sporophore branch architecture or shape
pinnate +
Sporophore branch quantity
2 +
Sporophore height
high +
Sporophore length or size
long +
Sporophore orientation
arising +
Sporophore position relational
level +
Sporophore presence
absent +
Sporophore quantity
2 +
Sporophore some measurement
0.5 cm5 mm <br />0.005 m <br /> (4 cm40 mm <br />0.04 m <br />) +
Sporophore structure in adjective form
sporangial +
Stalk height
high +
Stalk length
1/2 length of fertile segment +
Stalk quantity
common +
Stalk shape
flattened +
Stalk texture
not fleshy +
Stalk width
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
Stem architecture
unbranched +  and simple +
Stem orientation
upright +
Stem thickness
0 cm0 mm <br />0 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
Surface architecture or shape
baculate +
Surface arrangement
network +
Surface fragility
delicate +
Surface fusion
joined +
Surface pubescence or relief
warty +
Surface relief
tuberculate +  and rugate +
Taxon family
Ophioglossaceae +
Taxon name
Botrychium paradoxum +
Taxon parent
Botrychium subg. Botrychium +
Taxon rank
species +
Tissue architecture
vascular +
Trophophore architecture
sessile +  and short-stalked +
Trophophore blade architecture
Compound (Simple) +
Trophophore blade presence
absent +
Trophophore orientation
arising +
Vein architecture
anastomosing +
Vein arrangement
arranged +
Vein fusion
free +
Vein shape
pinnate +  and fan +
Volume
Volume 2 +
Whole organism growth form
plant +
Whole organism growth form or habitat
terrestrial +
Whole organism some measurement
0cm (15cm) +
X chromosome quantity
45 +