Botrychium sect. Hiemobotrychium

W. H. Wagner

Novon 2: 267. 1992.

Treatment appears in FNA Volume 2.

Roots 20–30, yellow to brown, 0.6–1 mm diam. 1cm from base. Plants over 6 cm. Common stalk with tracheidal idioblasts. Trophophore short-stalked and prostrate, arising from basal portion of common stalk near ground level; blades usually 2 per plant, appearing in late fall and dying in early spring, deltate, 2–3-pinnate, mostly 5–8 cm wide when mature, herbaceous. Leaf primordia with scattered hairs, hairs 0.6–1 mm. Leaf-sheath closed. Pinna divisions reduced to tip. First pinnule on basal pinnae usually borne basiscopically. Sporophores long-stalked, arising from basal portion of common stalk near ground level, commonly absent without primordia; stalks and rachis broadly flattened, fleshy, to 1.4 mm wide. x =45.

Distribution

Endemic to se United States

Discussion

Tracheidal idioblasts in Botrychium sect. Hiemobotrychium are enormous, elongate, spindle-shaped cells with helical or annular thickenings and are scattered along the vascular strand. They are unique among ferns.

Species 1.

Selected References

None.

Acute (?) +  and Rounded (?) +
Warren H. Wagner Jr. +  and Florence S. Wagner +
W. H. Wagner +
simple +  and 5-pinnate +
4 cm40 mm <br />0.04 m <br /> (25 cm250 mm <br />0.25 m <br />) +
deltate +
5cm;8cm +
Endemic to se United States +
0.1 cm1 mm <br />0.001 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
0.1 cm1 mm <br />0.001 m <br /> (1 cm10 mm <br />0.01 m <br />) +
0.06 cm0.6 mm <br />6.0e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
of widely scattered , long , uniseriate hairs +
dentate;lacerate +
spreading;ascending +
epiphytic +  and terrestrial +
spore-bearing +  and laminate +
photosynthetic +
with long , uniseriate hairs +  and glabrous +
flattened +
0 cm0 mm <br />0 m <br /> (0.14 cm1.4 mm <br />0.0014 m <br />) +
with corky ridges +  and smooth +
yellow +  and brown +
0.06 cm0.6 mm <br />6.0e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
not proliferous +
sessile +  and short-stalked +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
thick-walled +  and trilete +
flattened +
0 cm0 mm <br />0 m <br /> (0.14 cm1.4 mm <br />0.0014 m <br />) +
unbranched +  and simple +
0 cm0 mm <br />0 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
tuberculate +  and rugate +
Botrychium sect. Hiemobotrychium +
Botrychium subg. Sceptridium +
section +
anastomosing +
Pinnate (?) +  and Fan (?) +