Brachytheciastrum collinum

(Schleicher ex Müller Hal.) Ignatov & Huttunen

Arctoa 11: 260. 2003.

Illustrated
Basionym: Hypnum collinum Schleicher ex Müller Hal. Syn. Musc. Frond. 2: 429. 1851
Synonyms: Brachythecium collinum (Schleicher ex Müller Hal.) Schimper B. collinum var. idahense (Renauld & Cardot) Grout B. idahense Lesquereux H. hillebrandii
Treatment appears in FNA Volume 28. Treatment on page 410. Mentioned on page 409, 414, 461.

Plants small, in loose to dense tufts, green usually only when very young, pale-yellow or brownish yellow with age. Stems 2–4 cm, creeping to ascending, in dense tufts erect, terete-foliate, julaceous or subjulaceous, irregularly pinnate, branches to 4 mm, terete-foliate. Stem-leaves appressed proximally, closely imbricate, straight to falcate, ovate to broadly ovatelanceolate, not to rarely moderately plicate, 0.5–1 (–1.5) × 0.2–0.5 (–0.7) mm; margins plane or recurved proximally, coarsely serrate almost to base; apex abruptly or rarely gradually short-acuminate, acumen somewhat spreading to reflexed, often homomallous to falcate-secund; costa to 40–70% leaf length, occasionally almost absent, terminal spine absent; alar cells subquadrate, 8–20 × 9–11 µm, region small, of 10 × 4–6 cells, distinctly delimited; laminal cells elongate to linear, 25–60 × 4–7 µm; basal-cells almost undifferentiated. Seta reddish orange, 0.6–1.4 cm, slightly roughened, sometimes almost smooth except basal 1/4. Capsule slightly inclined to horizontal, reddish orange, ovate to subcylindric, 1–1.8 mm; endostome basal membrane 1/3 endostome length or longer, cilia as long as segments. Spores 9–12 µm.


Habitat: Soil, rock, xeric to mesic sunny to moderately shaded habitats
Elevation: low to high elevations (0-3700 m)

Distribution

V28 635-distribution-map.gif

Greenland, Alta., B.C., Sask., Yukon, Ariz., Calif., Colo., Idaho, Kans., Mont., Nebr., Nev., N.Mex., N.Dak., Oreg., S.Dak., Tex., Utah, Wash., Wyo., Mexico (Baja California), Mexico (Chihuahua), Europe, Asia, n Africa

Discussion

Although stems of Brachytheciastrum collinum are usually julaceous, the leaves are sometimes well spaced, making some populations difficult to distinguish from Brachythecium bolanderi. In this case, the rather extensive alar group and the autoicous sexual condition of B. collinum are helpful, as well as the more abruptly short-acuminate leaves. Some authors distinguish B. collinum and B. fendleri by the non-plicate leaves of the former. However, some collections, for example New Mexico, Santa Fe County, Buck 39717, have plicate leaves, while leaves of B. fendleri are usually smooth.

Selected References

None.

Lower Taxa

None.

"long" is not a number."broad" is not a number.

falcate-secund +
spreading;reflexed +
Michael S. Ignatov +
(Schleicher ex Müller Hal.) Ignatov & Huttunen +
undifferentiated +
slightly broader +
short-decurrent +
rounded +  and narrowed +
rounded-cucullate +  and decurrent +
short to long +
Hypnum collinum +
terete-foliate +
usually straight;curved +
cucullate +
reddish orange +
slightly inclined +  and horizontal +
ovate;subcylindric +
0.1 cm1 mm <br />0.001 m <br /> (0.18 cm1.8 mm <br />0.0018 m <br />) +
specialized +
synoicous +, dioicous +, autoicous +, sexual +  and asexual +
subquadrate +
differentiated +
9um +  and 11um +
3 (?) +  and 2 (?) +
discernible +
thick +  and thin +
autoicous +  and sexual +
Greenland +, Alta. +, B.C. +, Sask. +, Yukon +, Ariz. +, Calif. +, Colo. +, Idaho +, Kans. +, Mont. +, Nebr. +, Nev. +, N.Mex. +, N.Dak. +, Oreg. +, S.Dak. +, Tex. +, Utah +, Wash. +, Wyo. +, Mexico (Baja California) +, Mexico (Chihuahua) +, Europe +, Asia +  and n Africa +
yellow-green +  and orange tinge +
low to high elevations (0-3700 m) +
not adherent +
Soil, rock, xeric to mesic sunny to moderately shaded habitats +
4um (?) +  and 7um (?) +
pointed +
short-conic +
perfect +  and xerocastique +
hygrocastique +
opaque +  and clear +
reddish orange +
roughened +  and smooth +
0.6 cm6 mm <br />0.006 m <br /> (1.4 cm14 mm <br />0.014 m <br />) +
Illustrated +
0.1 cm1 mm <br />0.001 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.07 cm0.7 mm <br />7.0e-4 m <br />) +
erectopatent +  and erect +
ovate +  and broadly ovatelanceolate +
0.2mm;0.5mm +
terete-foliate +
pinnate +, subjulaceous +  and julaceous +
creeping +  and ascending +
2 cm20 mm <br />0.02 m <br /> (4 cm40 mm <br />0.04 m <br />) +
Brachythecium collinum +, B. collinum var. idahense +, B. idahense +  and H. hillebrandii +
Brachytheciastrum collinum +
Brachytheciastrum +
species +
incrassate +
thick;thick +
brownish yellow +, pale-yellow +  and green +