Braya humilis subsp. humilis

Illustrated
Synonyms: Arabidopsis novae-angliae (Rydberg) Britton Braya humilis var. abbei (Böcher) B. Boivin Braya humilis subsp. arctica (Rydberg) Hultén Braya humilis var. arctica (Böcher) B. Boivin Braya humilis var. interior (Böcher) B. Boivin Braya humilis var. laurentiana (Böcher) B. Boivin Braya humilis var. leiocarpa (Trautvetter) Fernald Braya humilis var. novae-angliae (Rydberg) Fernald Braya humilis subsp. richardsonii Rollins Braya humilis subsp. ventosa Böcher Braya humilis var. ventosa (Rollins) B. Boivin Braya intermedia Braya novae-angliae Braya novae-angliae subsp. abbei (Rollins) Böcher Braya novae-angliae var. interior Böcher Braya novae-angliae var. laurentiana Böcher Braya novae-angliae subsp. ventosa Böcher Braya richardsonii Pilosella novae-angliae Pilosella richardsonii Torularia humilis subsp. arctica
Treatment appears in FNA Volume 7. Treatment on page 549. Mentioned on page 550.

Stems ascending to erect, branched or unbranched, 0.4–3.3 dm, sparsely to moderately pubescent. Leaves: blade margins sinuate-dentate, shallowly pinnatifid, or entire, surfaces moderately pubescent. Flowers: petals white, pink, or purple, 2.5–6.9 (–7.5) × (0.7–) 0.9–4 (–4.2) mm. Fruits usually fertile and fully developed, somewhat torulose, 0.6–1.2 (–1.3) mm wide; septum not fenestrate or split longitudinally. 2n = 28, 42, 56, 70.


Phenology: Flowering May–Jul.
Habitat: Sandy, gravelly soil along streams, lakeshores, roadsides, moraines, open stony slopes, dolomite cliffs and slopes, limestone ledges, solifluction soils
Elevation: 0-4000 m

Distribution

V7 875-distribution-map.gif

Greenland, Alta., B.C., Man., Nfld. and Labr. (Nfld.), N.W.T., Nunavut, Ont., Que., Yukon, Alaska, Colo., Mich., Mont., Vt., Wyo., e, c Asia

Discussion

Subspecies humilis is extremely variable morphologically. In a general way, morphological form correlates with ploidy level, e.g., tetraploids, octoploids, and decaploids tend to be short in stature with small leaves. Hexaploids are less predictable. They range from short plants with small leaves to large, robust, multi-branched plants with large, pinnatifid leaves. Attempting to segregate most morphological forms of Braya humilis into logical infraspecific taxa is an exercise in futility. Populations that appear distinctive in the field almost always blur imperceptibly into the larger subsp. humilis continuum when compared with other populations from across the range of distribution. Subspecies humilis is broadly distributed on calcareous substrates in arctic, subarctic, alpine, and boreal regions of North America and Asia.

Selected References

None.

Lower Taxa

None.

"not" is not a number. "elongated" is not a number."thick" is not a number."dm" is not declared as a valid unit of measurement for this property."dm" is not declared as a valid unit of measurement for this property.

70 +, 56 +, 42 +  and 28 +
0.04 cm0.4 mm <br />4.0e-4 m <br /> (0.07 cm0.7 mm <br />7.0e-4 m <br />) +
obtuse +  and acute +
James G. Harris +
(C. A. Meyer) B. L. Robinson in A. Gray et al. +
cuneate +  and attenuate +
Sisymbrium humile +
appendaged +  and unappendaged +
2 cm20 mm <br />0.02 m <br /> (3.5 cm35 mm <br />0.035 m <br />) +
0.8 cm8 mm <br />0.008 m <br /> (1 cm10 mm <br />0.01 m <br />) +
not auriculate +
0.5 cm5 mm <br />0.005 m <br /> (2 cm20 mm <br />0.02 m <br />) +
entire +, pinnatifid +  and sinuate-dentate +
sublinear;oblong;sublinear;oblong;oblanceolate;spatulate;obovate +
smaller +
0.1 cm1 mm <br />0.001 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
urceolate +, campanulate +  and tubular +
branched +  and simple +
well-developed +
not +  and distinct +
not +  and differentiated +
emarginate +  and entire +
incumbent +  and accumbent +
subsessile +  and sessile +
0.8 cm8 mm <br />0.008 m <br /> (1.2 cm12 mm <br />0.012 m <br />) +
ascending +  and erect +
0.3 cm3 mm <br />0.003 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
Greenland +, Alta. +, B.C. +, Man. +, Nfld. and Labr. (Nfld.) +, N.W.T. +, Nunavut +, Ont. +, Que. +, Yukon +, Alaska +, Colo. +, Mich. +, Mont. +, Vt. +, Wyo. +, e +  and c Asia +
0-4000 m +
straight +  and curved +
not winged +  and unappendaged +
0.3 cm3 mm <br />0.003 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
connate +  and distinct +
not basally +  and dilated +
0.2 cm2 mm <br />0.002 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
actinomorphic +
latiseptate +  and sessile +
2.5 cm25 mm <br />0.025 m <br /> (3.2 cm32 mm <br />0.032 m <br />) +
0.18 cm1.8 mm <br />0.0018 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
1.2 cm12 mm <br />0.012 m <br /> (2.5 cm25 mm <br />0.025 m <br />) +
torulose +
0.06 cm0.6 mm <br />6.0e-4 m <br /> (0.12 cm1.2 mm <br />0.0012 m <br />) +
Sandy, gravelly soil along streams, lakeshores, roadsides, moraines, open stony slopes, dolomite cliffs and slopes, limestone ledges, solifluction soils +
sessile +  and petiolate +
alternate +, not +  and rosulate +
cauline +  and basal +
connate +  and distinct +
decurrent +
pinnatifid;sinuate-dentate;pinnatifid;sinuate-dentate +
tenuinucellate +, crassinucellate +  and bitegmic +
campylotropous +  and anatropous +
0.69 cm6.9 mm <br />0.0069 m <br /> (0.75 cm7.5 mm <br />0.0075 m <br />) +
0.4 cm4 mm <br />0.004 m <br /> (0.42 cm4.2 mm <br />0.0042 m <br />) +
purple +, pink +  and white +
2.5mm;6.9mm +
rudimentary +
spatulate +, oblanceolate +  and obovate +
0.09 cm0.9 mm <br />9.0e-4 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
Flowering May–Jul. +
trinucleate +  and 3(-11)-colpate +
in A. Gray et al., Syn. Fl. N. Amer. +
ascending +  and erect +
slender +
mucilaginous +
uniseriate +  and biseriate +
not +  and mucilaginous +
white +, black +, brown +  and yellow +
0.06 cm0.6 mm <br />6.0e-4 m <br /> (0.09 cm0.9 mm <br />9.0e-4 m <br />) +
0.04 cm0.4 mm <br />4.0e-4 m <br /> (0.05 cm0.5 mm <br />5.0e-4 m <br />) +
persistent +  and caducous +
distinct +
0.2 cm2 mm <br />0.002 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
0.08 cm0.8 mm <br />8.0e-4 m <br /> (0.12 cm1.2 mm <br />0.0012 m <br />) +
not fenestrate +
reduced +
latiseptate +  and sessile +
terete +, torulose +, globose +, lanceoloid-subulate +, lanceoloid +, ovoid +, oval-elliptic +, cylindrical +, oblong +  and linear +
latiseptate +  and sessile +
terete +, torulose +, globose +, lanceoloid-subulate +, lanceoloid +, ovoid +, oval-elliptic +, cylindrical +, oblong +  and linear +
Illustrated +
tetradynamous +
unbranched +  and branched +
procumbent +  and decumbent +
ascending +  and erect +
usually few +  and several +
2-lobed +  and entire +
subsessile +, sessile +  and petiolate +
0.08 cm0.8 mm <br />8.0e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
persistent +
distinct +
0.03 cm0.3 mm <br />3.0e-4 m <br /> (0.08 cm0.8 mm <br />8.0e-4 m <br />) +
Arabidopsis novae-angliae +, Braya humilis var. abbei +, Braya humilis subsp. arctica +, Braya humilis var. arctica +, Braya humilis var. interior +, Braya humilis var. laurentiana +, Braya humilis var. leiocarpa +, Braya humilis var. novae-angliae +, Braya humilis subsp. richardsonii +, Braya humilis subsp. ventosa +, Braya humilis var. ventosa +, Braya intermedia +, Braya novae-angliae +, Braya novae-angliae subsp. abbei +, Braya novae-angliae var. interior +, Braya novae-angliae var. laurentiana +, Braya novae-angliae subsp. ventosa +, Braya richardsonii +, Pilosella novae-angliae +, Pilosella richardsonii +  and Torularia humilis subsp. arctica +
Braya humilis subsp. humilis +
Braya humilis +
subspecies +
subsessile +  and short-stalked +
coiled +, 1-7-veined +  and veined +
glabrescent +, , +  and pubescent +
anastomosing +
not +  and scapose +
glabrous +, pilose +  and pubescent +
perennial +, biennial +  and annual +
aquatics +  and terrestrial +
glabrescent +  and pubescent +