Bucklandiella afoninae

(Frisvoll) Bednarek-Ochyra & Ochyra in R. Ochyra et al.

in R. Ochyra et al., Cens. Cat. Polish Mosses, 144. 2003,.

Basionym: Racomitrium afoninae Frisvoll J. Bryol. 15: 275, figs. 1, 2. 1988
Treatment appears in FNA Volume 27. Treatment on page 272. Mentioned on page 269, 271.

Plants fairly large, rather coarse, densely caespitose, olive green distally, brownish or blackish proximally. Stems erect-ascending, 2–5 cm, moderately, irregularly branched. Leaves loosely appressed, spirally twisted around the stem or somewhat contorted when dry, erect-spreading when moist, lanceolate, (2–) 2.3–2.7 (–2.9) × 0.5–0.6 mm, muticous or with a short, to 0.4 mm, hyaline, bluntly denticulate to crenulate hairpoint; margins broadly recurved to midleaf or 3/4 way up the leaf on one side, plane or narrowly recurved in the broadest part on the other side, variously 1–2-stratose in 1 cell row distally; costa percurrent, 80–90 µm wide basally, 55–75 µm wide distally, strongly convex on the abaxial side, 3-stratose in the base with 4 (–5) enlarged adaxial cells, 3–4-stratose in the middle with 3–4 large adaxial cells and 2–3-stratose distally with 2 adaxial cells; laminal cells 1-stratose with occasional 2-stratose patched near the apex, smooth or slightly pseudopapillose; distal and median cells rectangular, 25–50 × 10–12 µm, strongly sinuose-nodulose and thick-walled; basal-cells long-rectangular to linear, 50–90 × 10–12 µm, with strongly porose and incrassate lateral walls, to 7 µm thick; alar cells undifferentiated; basal marginal cells esinuose, moderately thick-walled, long-rectangular to linear, 50–100 × 5–8 µm, hyaline, forming a transparent border extending to the broadest part of the leaf. Sterile.


Habitat: Rocky ground in tundra

Discussion

Of conservation concern.

Bucklandiella afoninae is the rarest species of the genus, known in the flora area only from a single site from western Alaska (Seaward Peninsula), which corresponds to several localities of the species in Chukotka in arctic Russia. Although it is known only in the barren state, it fits well into the concept of sect. Marginatae on account of its linear basal cells with strongly incrassate and porose longitudinal walls, and the presence of a distinct basal marginal border that is composed of exceptionally long, mostly linear cells. Such leaf border is unknown in any other species of the genus in the flora area. The other exceptional character of this species is spirally twisted leaves, and such leaf arrangement is known only in two austral species of Bucklandiella, B. emersa and B. rupestris.

Selected References

None.

Lower Taxa

None.

"/2-3/4" is not declared as a valid unit of measurement for this property.

... more about "Bucklandiella afoninae"
large +  and enlarged +
sometimes +  and short-hyaline-tipped +
hyaline +  and yellowish hyaline +
papillose +
Ryszard Ochyra +  and Halina Bednarek-Ochyra +
(Frisvoll) Bednarek-Ochyra & Ochyra in R. Ochyra et al. +
both +  and papillose +
toothed +
nodulose;oblate;quadrate;elongate +
thick-walled +  and nodulose-porose +
long-rectangular;linear +
thick-walled +  and esinuose +
long-rectangular +  and linear +
divided +
Racomitrium afoninae +
transparent +
undifferentiated +
0 +  and 1/2 +
4-5-lobed +
small;large +
sulcate +  and smooth +
blackish brown +, castaneous +  and brown +
striate +  and smooth +
narrowed;obloid;narrowly cylindric +
transparent +
undifferentiated +
2-3-stratose;3-4-stratose;3-stratose;percurrent +
subpercurrent +
papillose +  and smooth +
55um +  and 75um +
sinuose-nodulose +  and rectangular +
Alaska +  and n Asia (Chukotka) +
sinuose-nodulose +
subquadrate +  and oblong +
Rocky ground in tundra +
0.27 cm2.7 mm <br />0.0027 m <br /> (0.29 cm2.9 mm <br />0.0029 m <br />) +
0.23 cm2.3 mm <br />0.0023 m <br /> (0.27 cm2.7 mm <br />0.0027 m <br />) +
erect-spreading +
muticous +, lanceolate +, bluntly denticulate +  and crenulate +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.06 cm0.6 mm <br />6.0e-4 m <br />) +
often entirely hyaline +  and yellowish hyaline +
pseudopapillose +  and smooth +
thick-walled +  and sinuose +
dioicous +  and sexual +
straight +
muticous +, piliferous +, falcate +  and lanceolate +
extensive +
indistinct +
long-rostellate;conic-rostrate +
not +  and differentiated +
lanceolate;linear +
brown +  and reddish-brown +
2-fid +, lanceolate +  and triangular +
2 +  and 3 +
terete +  and filiform +
in R. Ochyra et al., Cens. Cat. Polish Mosses, +
arcuate +  and straight +
papillose +  and smooth +
short to long +
ornamented +  and smooth +
spheric +
creeping +  and decumbent +
erect-ascending +
2 cm20 mm <br />0.02 m <br /> (5 cm50 mm <br />0.05 m <br />) +
Dryptodon subg. Ellipticodryptodon +, Racomitrium sect. Ellipticodryptodon +, Racomitrium subg. Ellipticodryptodon +  and Racomitrium subg. Microcarpa +
Bucklandiella afoninae +
Bucklandiella sect. Marginatae +
species +
compact +  and loose +
flexuose +  and straight +
incrassate +
thin +  and thick +
blackish;brownish;olive green +
plant +  and caespitose +