Bucklandiella sect. Sudeticae

(Bednarek-Ochyra) Bednarek-Ochyra & Ochyra in R. Ochyra et al.

in R. Ochyra et al., Cens. Cat. Polish Mosses, 143. 2003,.

Basionym: Racomitrium sect. Sudetica Bednarek-Ochyra Fragm. Florist. Geobot., Ser. Polon. 2: 221. 1995
Treatment appears in FNA Volume 27. Treatment on page 280. Mentioned on page 268.

Plants small, medium-sized to large, fairly slender to coarse and rigid. Stems 1–13 cm long, prostrate, ascending to erect, sparsely irregularly ramified to almost unbranched. Leaves lanceolate or ovatelanceolate, straight to curved; margins recurved on both sides to 1/2–3/4 the leaf length, sometimes to the apex, variously 1–2 (–4) -stratose in 1–3 (–6) cell rows; muticous or piliferous, awns erect or erect-recurved, not or slightly flexuose, stout, terete, strongly or moderately denticulate or spinulose-denticulate, not or slightly decurrent; costa 3–4-stratose and strongly convex abaxially in the proximal portion; alar cells undifferentiated; basal marginal border present. Inner perichaetial leaves slightly differentiated, hyaline and sheathing in the proximal portion, chlorophyllose distally, usually piliferous and with the areolation similar to that in the vegetative leaves. Peristome teeth with a basal membrane.

Distribution

North America, South America, Arctic and temperate Asia, Atlantic Islands, Australia, Antarctica

Discussion

Species 4 (4 in the flora).

The slightly differentiated innermost perichaetial bracts that are ovate-lanceolate and similar to the vegetative leaves, the stout, terete or somewhat flattened, not flexuose and variously denticulate and spinulose awn, presence of the basal marginal border, laminal cells tending to be short or very short in the distal portion, and strong, 3–4-stratose costa in the basal part are diagnostic of sect. Sudeticae.

Selected References

None.

Key

1 Costa canaliculate, very broad, 100-120 µm, with 6-9 adaxial cells in the basal part; laminal cells strongly pseudopapillose; awns broad and decurrent, coarsely denticulate and spinulose. Bucklandiella brevipes
1 Costa carinate, narrower, 60-100 µm, with 3-4(-5) adaxial cells in the basal part; laminal cells not or weakly pseudopapillose; awns not decurrent, less coarsely papillose and denticulate > 2
2 Costa shallowly grooved and winged on the abaxial side, asymmetric in outline; leaf margins lumpy distally; awns present, terete, sharply spinulose. Bucklandiella occidentalis
2 Costa symmetric in outline, not furrowed and winged abaxially; leaf margins smooth, awns present or absent, not terete, weakly and obtusely denticulate and spinulose > 3
3 Leaf margins regularly 2(-3)-stratose for 2-4(-6) cell rows; costa predominantly 4-stratose basally; awns, if present, short, to 0.2 mm, yellowish or reddish hyaline; plants mostly reddish brown Bucklandiella macounii
3 Leaf margins 1-2-stratose for 1 cell row or occasionally 2(-3)-stratose for 2(-3) cell rows; costa predominantly (2-)3(-4)-stratose basally; awns present, longer than 0.2 mm, hyaline; plants never reddish brown. Bucklandiella sudetica
sometimes +  and short-hyaline-tipped +
hyaline +  and yellowish hyaline +
papillose +
Ryszard Ochyra +  and Halina Bednarek-Ochyra +
(Bednarek-Ochyra) Bednarek-Ochyra & Ochyra in R. Ochyra et al. +
flexuose +
erect-recurved +  and erect +
both +  and papillose +
decurrent +, spinulose-denticulate +, denticulate +, or +  and terete +
nodulose;oblate;quadrate;elongate +
thick-walled +  and nodulose-porose +
rectangular;linear +
divided +
Racomitrium sect. Sudetica +
undifferentiated +
0 +  and 1/2 +
4-5-lobed +
small;large +
sulcate +  and smooth +
blackish brown +, castaneous +  and brown +
striate +  and smooth +
narrowed;obloid;narrowly cylindric +
undifferentiated +
1 (?) +  and 3 (?) +
3-4-stratose +
subpercurrent +
papillose +  and smooth +
North America +, South America +, Arctic and temperate Asia +, Atlantic Islands +, Australia +  and Antarctica +
sinuose-nodulose +
subquadrate +  and oblong +
sinuose +, multistratose +  and 1-stratose +
pseudopapillose +
thick-walled +  and sinuose +
dioicous +  and sexual +
straight +  and curved +
1-2(-4)-stratose +
piliferous +, muticous +, ovatelanceolate +  and lanceolate +
1-stratose;variously multistratose +
cristate +  and serrate +
extensive +
indistinct +
long-rostellate;conic-rostrate +
not +  and differentiated +
lanceolate;linear +
brown +  and reddish-brown +
2-fid +, lanceolate +  and triangular +
2 +  and 3 +
terete +  and filiform +
in R. Ochyra et al., Cens. Cat. Polish Mosses, +
arcuate +  and straight +
papillose +  and smooth +
short to long +
0 +  and 1/2-3/4 +
ornamented +  and smooth +
spheric +
unbranched +, ramified to almost +  and ramified +
creeping +  and decumbent +
1cm;13cm +
ascending +  and erect +
Dryptodon subg. Ellipticodryptodon +, Racomitrium sect. Ellipticodryptodon +, Racomitrium subg. Ellipticodryptodon +  and Racomitrium subg. Microcarpa +
Bucklandiella sect. Sudeticae +
Bucklandiella +
section +
compact +  and loose +
flexuose +  and straight +
thin +  and thick +
jet-black;blackish brown;olive green;yellowish;brown;green +
slender;medium-sized;large +