Camissoniopsis bistorta

(Nuttall ex Torrey & A. Gray) W. L. Wagner & Hoch

Syst. Bot. Monogr. 83: 204. 2007.

Basionym: Oenothera bistorta Nuttall ex Torrey & A. Gray Fl. N. Amer. 1: 508. 1840
Synonyms: Camissonia bistorta (Nuttall ex Torrey & A. Gray) P. H. Raven O. bistorta var. veitchiana Hooker Sphaerostigma bistortum (Nuttall ex Torrey & A. Gray) Walpers S. bistortum var. veitchianum (Hooker) A. Nelson S. veitchianum (Hooker) Small
Treatment appears in FNA Volume 10.

Herbs annual, rarely short-lived perennial, usually villous, sometimes strigillose. Stems 1–several from base, ascending or decumbent, to 80 cm. Leaves 1.2–12 × 0.2–1.5 cm; petiole 0–4 cm, distal ones 0–0.3 cm; blade (basal) narrowly elliptic or (cauline) usually narrowly lanceolate or lanceolate, rarely linear, base (basal) narrowly cuneate, (cauline) cuneate or subcordate, margins usually sparsely and incon­spicuously denticulate, apex acute. Flowers opening near sunrise; floral-tube 2–5 (–7.5) mm; sepals (2.3–) 5–8 (–11) mm; petals yellow, each usually with 1 bright red dot, rarely 2, near base, (4.2–) 7–15 mm; epise­palous filaments (1–) 1.5–3.5 mm, epipetalous filaments (0.5–) 1–2.5 mm, anthers (0.5–) 1.3–2 (–2.5) mm, less than 5% of pollen-grains 4-pored or 5-pored; style (5.5–) 7–12 mm, stigma exserted beyond anthers at anthesis. Capsules straight or somewhat contorted, weakly 4-angled, 12–40 × 1.5–2.5 mm. Seeds 0.9–1 mm. 2n = 14.


Phenology: Flowering Mar–Jun.
Habitat: Sandy or clayey soils, coastal strands, grasslands, coastal sage scrub, chaparral, oak woodlands, margins of Sonoran and Mojave deserts, rarely higher elevation meadows.
Elevation: 0–1600(–2600) m.

Distribution

Calif., Mexico (Baja California)

Discussion

Camissoniopsis bistorta occurs in California from Ventura County south and east through the counties of southern Los Angeles, southwestern San Bernardino, Orange, western Riverside, and the western two-thirds of San Diego, reaching the margins of the desert in San Bernardino and San Diego counties, and southward in cismontane Baja California to Ojos Negros and San Vicente. The species occurs at exceptionally high elevations in the Santa Ana drainage of the San Bernardino Mountains. P. H. Raven (1969) indicated that there were occasional apparent hybrids between C. cheiranthifolia subsp. suffruticosa and C. bistorta occurring in intermediate habitats in areas where the two species co-occur. He determined that C. bistorta is self-incompatible.

Camissoniopsis bistorta was apparently introduced with stream gravel in 1959 in Goleta Marsh, Santa Barbara, California, and on ballast heaps at Nanaimo, Vancouver Island, British Columbia, in 1893. It has apparently not persisted at either site.

Oenothera heterophylla Nuttall ex Hooker & Arnott (1839), not Spach (1836), is an illegitimate name that pertains to Camissoniopsis bistorta.

Selected References

None.

Lower Taxa

None.
... more about "Camissoniopsis bistorta"
0.2 cm2 mm <br />0.002 m <br /> (0.25 cm2.5 mm <br />0.0025 m <br />) +
0.13 cm1.3 mm <br />0.0013 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
Warren L. Wagner +
(Nuttall ex Torrey & A. Gray) W. L. Wagner & Hoch +
subcordate +  and cuneate +
Oenothera bistorta +
indehiscent +  and loculicidal +
serrulate +, denticulate +  and dentate +
lanceolate +  and elliptic +
1.2 cm12 mm <br />0.012 m <br /> (4 cm40 mm <br />0.04 m <br />) +
1-5 times or straight narrowly cylindrical and thickened +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.25 cm2.5 mm <br />0.0025 m <br />) +
Calif. +  and Mexico (Baja California) +
bright red +
2 +  and 1 +
0–1600(–2600) m. +
white +  and reddish-brown +
exfoliating +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
0.1 cm1 mm <br />0.001 m <br /> (0.25 cm2.5 mm <br />0.0025 m <br />) +
0.1 cm1 mm <br />0.001 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
basifixed +  and versatile +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.35 cm3.5 mm <br />0.0035 m <br />) +
0.5 cm5 mm <br />0.005 m <br /> (0.75 cm7.5 mm <br />0.0075 m <br />) +
0.2 cm2 mm <br />0.002 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
actinomorphic +
not swollen +, 4-angled +, curled +  and contorted +
1-5 times or straight narrowly cylindrical and thickened +
Sandy or clayey soils, coastal strands, grasslands, coastal sage scrub, chaparral, oak woodlands, margins of Sonoran and Mojave deserts, rarely higher elevation meadows. +
perennial +, short-lived +  and annual +
strigillose +  and villous +
deciduous +
1.2 cm12 mm <br />0.012 m <br /> (12 cm120 mm <br />0.12 m <br />) +
0.2 cm2 mm <br />0.002 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
denticulate +
numerous +  and 1 +
0.42 cm4.2 mm <br />0.0042 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
0.7 cm7 mm <br />0.007 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
0 cm0 mm <br />0 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
Flowering Mar–Jun. +
Syst. Bot. Monogr. +
basal +  and cauline +
clavate +  and globose +
7 +  and numerous +
sculptured +  and smooth +
obovoid +  and flattened +
0.09 cm0.9 mm <br />9.0e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
0.8 cm8 mm <br />0.008 m <br /> (1.1 cm11 mm <br />0.011 m <br />) +
purple +, pink +, red +  and green +
deciduous +
0.5 cm5 mm <br />0.005 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
parenchymatous +
parietal +, axile +  and placentation +
unequal +
aquatic +, amphibious +  and terrestrial +
2 times as many or as many as sepals +
prostrate +  and decumbent +
decumbent +  and ascending +
1 +  and several +
0 cm0 mm <br />0 m <br /> (80 cm800 mm <br />0.8 m <br />) +
subcapitate +  and subglobose +
petiolate +  and sessile +
intrapetiolar +
0.55 cm5.5 mm <br />0.0055 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
pubescent +  and glabrous +
0.7 cm7 mm <br />0.007 m <br /> (1.2 cm12 mm <br />0.012 m <br />) +
aquatic +, amphibious +  and terrestrial +
non-papillate +
Camissonia bistorta +, O. bistorta var. veitchiana +, Sphaerostigma bistortum +, S. bistortum var. veitchianum +  and S. veitchianum +
Camissoniopsis bistorta +
Camissoniopsis +
species +
3(-5)-aperturate +
smooth +  and erose +
papillate +