Camissoniopsis guadalupensis subsp. clementiana

(P. H. Raven) W. L. Wagner & Hoch

Syst. Bot. Monogr. 83: 204. 2007.

Basionym: Oenothera guadalupensis subsp. clementina P. H. Raven Aliso 5: 332. 1963
Synonyms: Camissonia guadalupensis subsp. clementina (P. H. Raven) P. H. Raven
Treatment appears in FNA Volume 10.

Herbs annual, subsucculent and heavy-set, densely villous, also glandular puberulent distally. Stems erect, with branches usually arising near base, 2–18 (–35) cm. Leaves 1.2–3.8 (–9.5) × 0.5–1.2 (–1.8) cm; petiole 0–3 cm; blade (basal and proximal cauline) narrowly elliptic, (cauline) narrowly ovate, base attenuate, margins sparsely and inconspicuously denticulate, apex (basal) acute, (cauline) obtuse, rounded, or truncate. Flowers opening near sunrise; floral-tube 1.6–2.4 mm; sepals 1.9–3.2 mm; petals yellow, each with 1 red dot near base, 2.8–4.2 mm; episepalous filaments 1.3–2.3 mm, epipetalous filaments 0.4–1.6 mm, anthers 0.4–0.8 mm, less than 5% of pollen-grains 4-pored or 5-pored; style 3.2–4.5 mm, stigma surrounded by anthers at anthesis. Capsules stout, straight or slightly curved outward, 4-angled, 10–18 × 2.8–3.5 mm, deeply grooved along lines of dehiscence. Seeds 0.7–0.9 mm. 2n = 14.


Phenology: Flowering Mar–Jun(–Sep).
Habitat: Sandy flats, dunes.
Elevation: 0–60 m.

Discussion

Subspecies clementiana is restricted to San Clemente Island, Los Angeles County, where it is common on dunes around the north end and down the west shore, perhaps to the south end.

Selected References

None.

Lower Taxa

None.
0.04 cm0.4 mm <br />4.0e-4 m <br /> (0.08 cm0.8 mm <br />8.0e-4 m <br />) +
truncate +, rounded +, obtuse +  and acute +
Warren L. Wagner +
(P. H. Raven) W. L. Wagner & Hoch +
attenuate +
Oenothera guadalupensis subsp. clementina +
indehiscent +  and loculicidal +
serrulate +, denticulate +  and dentate +
curved +  and straight +
1 cm10 mm <br />0.01 m <br /> (1.8 cm18 mm <br />0.018 m <br />) +
1-5 times or straight narrowly cylindrical and thickened +
0.28 cm2.8 mm <br />0.0028 m <br /> (0.35 cm3.5 mm <br />0.0035 m <br />) +
0–60 m. +
white +  and reddish-brown +
exfoliating +
0.04 cm0.4 mm <br />4.0e-4 m <br /> (0.16 cm1.6 mm <br />0.0016 m <br />) +
0.13 cm1.3 mm <br />0.0013 m <br /> (0.23 cm2.3 mm <br />0.0023 m <br />) +
basifixed +  and versatile +
0.16 cm1.6 mm <br />0.0016 m <br /> (0.24 cm2.4 mm <br />0.0024 m <br />) +
actinomorphic +
not swollen +, 4-angled +, curled +  and contorted +
1-5 times or straight narrowly cylindrical and thickened +
Sandy flats, dunes. +
puberulent +  and villous +
3.8 cm38 mm <br />0.038 m <br /> (9.5 cm95 mm <br />0.095 m <br />) +
1.2 cm12 mm <br />0.012 m <br /> (1.8 cm18 mm <br />0.018 m <br />) +
deciduous +
1.2 cm12 mm <br />0.012 m <br /> (3.8 cm38 mm <br />0.038 m <br />) +
0.5 cm5 mm <br />0.005 m <br /> (1.2 cm12 mm <br />0.012 m <br />) +
denticulate +
numerous +  and 1 +
0.28 cm2.8 mm <br />0.0028 m <br /> (0.42 cm4.2 mm <br />0.0042 m <br />) +
0 cm0 mm <br />0 m <br /> (3 cm30 mm <br />0.03 m <br />) +
Flowering Mar–Jun(–Sep). +
Syst. Bot. Monogr. +
basal +  and cauline +
clavate +  and globose +
7 +  and numerous +
sculptured +  and smooth +
obovoid +  and flattened +
0.07 cm0.7 mm <br />7.0e-4 m <br /> (0.09 cm0.9 mm <br />9.0e-4 m <br />) +
purple +, pink +, red +  and green +
deciduous +
0.19 cm1.9 mm <br />0.0019 m <br /> (0.32 cm3.2 mm <br />0.0032 m <br />) +
parenchymatous +
parietal +, axile +  and placentation +
unequal +
aquatic +, amphibious +  and terrestrial +
2 times as many or as many as sepals +
18 cm180 mm <br />0.18 m <br /> (35 cm350 mm <br />0.35 m <br />) +
prostrate +  and decumbent +
2 cm20 mm <br />0.02 m <br /> (18 cm180 mm <br />0.18 m <br />) +
subcapitate +  and subglobose +
petiolate +  and sessile +
intrapetiolar +
pubescent +  and glabrous +
0.32 cm3.2 mm <br />0.0032 m <br /> (0.45 cm4.5 mm <br />0.0045 m <br />) +
aquatic +, amphibious +  and terrestrial +
non-papillate +
Camissonia guadalupensis subsp. clementina +
Camissoniopsis guadalupensis subsp. clementiana +
Camissoniopsis guadalupensis +
subspecies +
3(-5)-aperturate +
smooth +  and erose +
papillate +