Castilleja affinis

Hooker & Arnott

Bot. Beechey Voy., 154. 1833.

Common names: California or coast paintbrush
Illustrated
Treatment appears in FNA Volume 17. Treatment on page 582. Mentioned on page 566, 572, 576, 577, 580, 581, 583, 584, 624, 665.

Herbs, perennial, 1.4–6 dm; caudex woody; with a taproot. Stems few-to-many, erect to ascending, unbranched or branched at proximal nodes, sometimes with short, leafy axillary branches, hairs sparse to dense, spreading, short and long, soft to stiff, unbranched, sometimes branched, eglandular, sometimes stipitate-glandular. Leaves green or purplish, sometimes redbrown, linear or linear-oblong to broadly lanceolate, 2–13 cm, not or ± fleshy, margins wavy or plane, flat or involute, 0–5-lobed, sometimes with small secondary lobes, short, leafy axillary shoots common, usually conspicuous, apex acuminate or acute to rounded; lobes spreading, linear to lanceolate, lateral lobes almost as wide as mid-blade, apex acute to rounded. Inflorescences 3–25 (–30) × 1.5–5 cm; bracts proximally green or deep purple, distally red, crimson, scarlet, pink, pinkish purple, pinkish red, or yellow, sometimes rose magenta, red-orange, or orange, oblanceolate or obovate to oblong or lanceolate, (0–) 3–5 (–7) -lobed; lobes spreading to ascending, linear to obovate, long, proximal lobes at or arising below mid length, apices acute to obtuse, center lobe sometimes rounded. Pedicels 0 mm or nearly so. Calyces proximally pale or green, distally as in distal portion of bracts, 14–35 mm; abaxial and adaxial clefts 6–22 mm, 33–50% of calyx length, deeper than laterals, lateral 2–7 mm, 15–25% of calyx length; lobes lanceolate or oblong, apex acute to obtuse or rounded, sometimes curved upward. Corollas straight or ± curved, 17–40 mm; tube 10–15 mm; beak long-exserted to subequal to calyx, adaxially green, 7–20 mm, puberulent, eglandular; abaxial lip deep green to reddish, brown, or deep purple, reduced, inconspicuous, included in calyx, 1.5–3 mm, 15–25% as long as beak; teeth ascending, green, 0.5–2 mm. 2n = 48, 72, 96.

Discussion

Varieties 3 (3 in the flora).

Castilleja affinis is highly variable and one of the more common paintbrushes at lower elevations in California, west of the Sierra Nevada, from the northern coast south to northern Baja California. Some recent authors (for example, M. Wetherwax et al. 2012) include C. litoralis of the Pacific Northwest coast as a subspecies of C. affinis, but due to the high polyploid nature of C. litoralis and its significantly closer morphological resemblance to C. miniata var. dixonii, that treatment is not followed here. Also see the comments under 62. C. litoralis.

Selected References

None.

Key

1 Stem hairs branched; distal portion of bracts usually pink to pinkish purple or pinkish red; coastal San Luis Obispo and Santa Barbara counties. Castilleja affinis var. contentiosa
1 Stem hairs unbranched, rarely branched; distal portion of bracts usually bright red, red-orange, crimson, scarlet, or yellow; more widespread in w California. > 2
2 Distal portions of bracts bright red to red-orange, crimson, or scarlet, rarely yellow, orange, rose, magenta, or pinkish red; inflorescences (2.5–)3–5 cm wide; chaparral slopes, openings, open woods, coastal scrub, stabilized dunes; much of w California. Castilleja affinis var. affinis
2 Distal portions of bracts yellow, rarely pink or pale red-orange, often becoming reddish tinted after anthesis; inflorescences 1.5–2.5 cm wide; serpentine substrates; San Francisco Bay region. Castilleja affinis var. neglecta

"acute" is not a number."nearly" is not a number.

... more about "Castilleja affinis"
96 +, 72 +  and 48 +
green +  and reddish brown or deep purple +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
J. Mark Egger +, Peter F. Zika +, Barbara L. Wilson +, Richard E. Brainerd +  and Nick Otting +
Hooker & Arnott +
long-exserted +
puberulent +
subequal +
0.7 cm7 mm <br />0.007 m <br /> (2 cm20 mm <br />0.02 m <br />) +
not +, leathery +  and fleshy +
orange;red-orange;orange;red-orange;rose morange;red-orange;orange;red-orange;rose magenta;yellow;pinkish red;pinkish purple;pink;scarlet;crimson;red;purple;deep;green;yellow;pinkish red;pinkish purple;pink;scarlet;crimson;red;purple;deep;green;yellow;pinkish red;pinkish purple;pink;scarlet;crimson;red;purple;deep;green;yellow;pinkish red;pinkish purple;pink;scarlet;crimson;red;purple;deep;green;yellow;pinkish red;pinkish purple;pink;scarlet;crimson;red;purple;deep;green;yellow;pinkish red;pinkish purple;pink;scarlet;crimson;red;purple;deep;green;yellow;pinkish red;pinkish purple;pink;scarlet;crimson;red;purple;deep;green;yellow;pinkish red;pinkish purple;pink;scarlet;crimson;red;purple;deep;greenpink;scarlet;crimson;red;purple;deep;green +
(0-)3-5(-7)-lobed +, obovate +  and oblong or lanceolate +
lateral +  and deeper +
0.2 cm2 mm <br />0.002 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
green +  and pale +
2-lobed +, lobed +  and tubular +
1.4 cm14 mm <br />0.014 m <br /> (3.5 cm35 mm <br />0.035 m <br />) +
taprooted;fibrous-rooted or rhizomatous +
California or coast paintbrush +
purple +, pink +, red +, orange +, yellow +, white +, green +  and pale greenish +
curved +  and straight +
1.7 cm17 mm <br />0.017 m <br /> (4 cm40 mm <br />0.04 m <br />) +
Calif. +  and nw Mexico +
spreading-hairy +  and glabrous +
2 +  and 1 +
indehiscent +, septicidal +, loculicidal +  and dehiscence +
eglandular +, branched +  and unbranched +
sparse +  and dense +
stipitate-glandular +
25 cm250 mm <br />0.25 m <br /> (30 cm300 mm <br />0.3 m <br />) +
axillary +  and terminal +
spiral +, whorled +, alternate +  and opposite +
redbrown +, purplish +  and green +
deciduous +
basal +  and cauline +
linear-oblong;broadly lanceolate +
2 cm20 mm <br />0.02 m <br /> (13 cm130 mm <br />0.13 m <br />) +
spreading +  and ascending +
3 +, 2 +  and 4 +
oblong +, lanceolate +, linear +  and obovate +
subequal +
0-5-lobed +, involute +, flat +, plane +  and wavy +
tenuinucellate +  and unitegmic +
campylotropous-like +
Bot. Beechey Voy., +
20 +  and 100 +
trapezoidal +, oblong-ovoid +, pyramidal +  and ovoid +
Illustrated +
non-secund +  and secund +
elongate +
subequal +
branched +  and unbranched +
sprawling +  and prostrate +
aerial +  and subterranean +
erect +  and ascending +
not fleshy +
2-lobed +  and entire +
Euchroma +  and Oncorhynchus +
Castilleja affinis +
Castilleja +
species +
shallow;deep +
reduced +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
1 cm10 mm <br />0.01 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +