Castilleja ambigua var. meadii

J. M. Egger & Ruygt

Phytoneuron 2012-68: 2, figs. 1, 3–7, 9[left]. 2012.

Common names: Mead’s owl’s-clover
EndemicConservation concern
Treatment appears in FNA Volume 17. Treatment on page 586. Mentioned on page 584.

Stems erect, unbranched, sometimes with a few divaricate-ascending branches from proximal 1/2 of stem above base. Leaves linear or linear-lanceolate to lanceolate, 1 mm wide at base, not fleshy, apex acuminate. Bracts proximally pale greenish, distally white on lobe apices, often becoming entirely greenish with age; lobes divaricate-ascending, linear, 8–14 mm, usually arising below mid length. Calyces with all 4 clefts subequal or lateral clefts shallower; lateral clefts 2–3 mm, 25% of calyx length. Corollas 14–21 mm; beak pale, off-white, pale-yellow, green with margins off-white, or yellow, sometimes orange, 1–4 mm; abaxial lip pale-yellow; teeth white to green.


Phenology: Flowering Apr–Jun(–Jul).
Habitat: Seasonally wet meadows with volcanic substrates in oak-pine woodlands or chaparral, shallow vernal pools, ephemeral stream margins.
Elevation: 400–500 m.

Discussion

Variety meadii is limited to vernally wet habitats, growing over rocks of the Sonoma Volcanic Formation in central Napa County near Atlas Peak. All known populations are under private ownership, and the variety is of conservation concern due to its very limited range. For the present, all populations except one are protected by conservation easements. Variety meadii often grows alongside, but does not hybridize with, Castilleja attenuata and C. densiflora. Ongoing study of annual species of Castilleja suggests this variety is genetically distinctive and may deserve full species status (S. J. Jacobs et al. 2018).

Selected References

None.

Lower Taxa

None.

0.2 cm2 mm <br />0.002 m <br /> (0.55 cm5.5 mm <br />0.0055 m <br />) +
stipitate-glandular +
J. Mark Egger +, Peter F. Zika +, Barbara L. Wilson +, Richard E. Brainerd +  and Nick Otting +
J. M. Egger & Ruygt +
whitish +, purple spots +  and redbrown +
0.1 cm1 mm <br />0.001 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
Orange (?) +, Yellow (?) +, Off-white (?) +, Green (?) +, Pale-yellow (?) +  and Pale (?) +
curved +  and straight +
short-hairy +
0.1 cm1 mm <br />0.001 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
not +, leathery +  and fleshy +
greenish;white;pale greenish +
(0-)3-5(-7)-lobed +, ovate +, oblong +  and lanceolate +
divaricate-ascending +
purple +, white +, green +  and pale yellowish +
2-lobed +, lobed +  and tubular +
0.8 cm8 mm <br />0.008 m <br /> (2.3 cm23 mm <br />0.023 m <br />) +
taprooted;fibrous-rooted or rhizomatous +
fleshy +  and woody +
subequal +
Mead’s owl’s-clover +
purple +, pink +, red +, orange +, yellow +, white +, green +  and pale greenish +
1.4 cm14 mm <br />0.014 m <br /> (2.1 cm21 mm <br />0.021 m <br />) +
400–500 m. +
2 +  and 1 +
indehiscent +, septicidal +, loculicidal +  and dehiscence +
Seasonally wet meadows with volcanic substrates in oak-pine woodlands or chaparral, shallow vernal pools, ephemeral stream margins. +
sparse +  and moderately dense +
short-stipitate-glandular +
9 cm90 mm <br />0.09 m <br /> (13 cm130 mm <br />0.13 m <br />) +
axillary +  and terminal +
spiral +, whorled +, alternate +  and opposite +
0.6 cm6 mm <br />0.006 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
brownish +  and green +
deciduous +
basal +  and cauline +
linear-lanceolate;lanceolate +
0.8 cm8 mm <br />0.008 m <br /> (5 cm50 mm <br />0.05 m <br />) +
not fleshy +
1 cm10 mm <br />0.01 m <br /> (?) +
arising +  and divaricate-ascending +
3 +, 2 +  and 4 +
narrowly lanceolate +  and oblong or more or less triangular +
subequal +
short to long +
yellow +  and off-white +
3-5(-7)-lobed +  and plane +
acute;obtuse +
tenuinucellate +  and unitegmic +
campylotropous-like +
Flowering Apr–Jun(–Jul). +
0.2 cm2 mm <br />0.002 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
0.3 cm3 mm <br />0.003 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
Phytoneuron +
20 +  and 100 +
trapezoidal +, oblong-ovoid +, pyramidal +  and ovoid +
Endemic +  and Conservation concern +
non-secund +  and secund +
elongate +
subequal +
aerial +  and subterranean +
not fleshy +
2-lobed +  and entire +
Euchroma +  and Oncorhynchus +
Castilleja ambigua var. meadii +
Castilleja ambigua +
variety +
white +  and green +
shallow;deep +
spreading +  and erect +
reduced +
0.1 cm1 mm <br />0.001 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
expanded +
1.1 cm11 mm <br />0.011 m <br /> (2.1 cm21 mm <br />0.021 m <br />) +
biennial +  and annual +