Castilleja applegatei var. breweri

(Fernald) N. H. Holmgren in A. Cronquist et al.

Intermount. Fl. 4: 486. 1984.

Common names: Brewer’s paintbrush
Endemic
Basionyms: Castilleja breweri Fernald Erythea 6: 49. 1898
Synonyms: C. applegatei subsp. pallida (Eastwood) T. I. Chuang & Heckard C. applegatei var. pallida (Eastwood) N. H. Holmgren C. breweri var. pallida Eastwood C. glandulifera subsp. pallida (Eastwood) Pennell
Treatment appears in FNA Volume 17. Treatment on page 588. Mentioned on page 589.

Herbs (0.8–) 1–3 (–4.8) dm. Stems branched from base. Leaves linear to broadly lanceolate, 1.2–4.8 cm, (0–) 3 (–5) -lobed, apex acute, rarely obtuse. Inflorescences 2–12 × 2–3.5 cm; bracts distally red, scarlet, or pale orange, rarely yellow, 3-lobed, apex acute to rounded, central lobe sometimes expanded. Calyces green to yellow, lobe apices red to yellow, 13–15 mm; abaxial and adaxial clefts (4–) 5–8 (–10) mm, 33% of calyx length, lateral 3–5 (–7) mm, 10–25% of calyx length; lobe apex acute, rarely obtuse. Corollas 16–22 mm; tube 11–13 mm; beak 6–10 mm. 2n = 48.


Phenology: Flowering Jun–Aug(–Sep).
Habitat: Dry rocky slopes, flats, open montane forests to alpine, granitic substrates.
Elevation: 1800–3600 m.

Discussion

Variety breweri is found in California in the high Sierra Nevada and in the mountains of adjacent Nevada. Hybrids with Castilleja chromosa and C. miniata var. miniata have both been found in Inyo County, California.

Selected References

None.

Lower Taxa

None.

0.5 cm5 mm <br />0.005 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
0.3 cm3 mm <br />0.003 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
green +  and whitish or yellow +
exserted +  and included +
protuberant +  and inconspicuous +
0.1 cm1 mm <br />0.001 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
Acute (?) +  and Rounded (?) +
J. Mark Egger +, Peter F. Zika +, Barbara L. Wilson +, Richard E. Brainerd +  and Nick Otting +
(Fernald) N. H. Holmgren in A. Cronquist et al. +
Castilleja breweri +
long-exserted +
0.6 cm6 mm <br />0.006 m <br /> (1 cm10 mm <br />0.01 m <br />) +
not +, leathery +  and fleshy +
yellow;pale orange;scarlet;pale orange;scarlet;red +
3-lobed +
green +  and yellow +
2-lobed +, lobed +  and tubular +
taprooted;fibrous-rooted or rhizomatous +
fleshy +  and woody +
Brewer’s paintbrush +
3.5 cm35 mm <br />0.035 m <br /> (4.1 cm41 mm <br />0.041 m <br />) +
purple +, pink +, red +, orange +, yellow +, white +, green +  and pale greenish +
curved +  and straight +
1.6 cm16 mm <br />0.016 m <br /> (2.2 cm22 mm <br />0.022 m <br />) +
Calif. +  and Nev. +
1800–3600 m. +
spreading-hairy +  and glabrous +
2 +  and 1 +
indehiscent +, septicidal +, loculicidal +  and dehiscence +
Dry rocky slopes, flats, open montane forests to alpine, granitic substrates. +
sparse +  and dense +
stipitate-glandular +
axillary +  and terminal +
spiral +, whorled +, alternate +  and opposite +
green +  and purplish or brown +
deciduous +
basal +  and cauline +
(0-)3(-5)-lobed;linear;broadly lanceolate +
1.2 cm12 mm <br />0.012 m <br /> (4.8 cm48 mm <br />0.048 m <br />) +
not fleshy +
1.3 cm13 mm <br />0.013 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
arising +, spreading +  and ascending +
3 +, 2 +  and 4 +
triangular +, oblong +, lanceolate-acuminate +, lanceolate +  and broadly lanceolate +
expanded +
(0-)3(-5)-lobed +  and wavy +
acuminate;obtuse;acuminate;obtuse;rounded;acute +
tenuinucellate +  and unitegmic +
campylotropous-like +
Flowering Jun–Aug(–Sep). +
Intermount. Fl. +
20 +  and 100 +
trapezoidal +, oblong-ovoid +, pyramidal +  and ovoid +
non-secund +  and secund +
elongate +
subequal +
sprawling +  and prostrate +
aerial +  and subterranean +
decumbent +, ascending +  and erect +
not fleshy +
2-lobed +  and entire +
C. applegatei subsp. pallida +, C. applegatei var. pallida +, C. breweri var. pallida +  and C. glandulifera subsp. pallida +
Castilleja applegatei var. breweri +
Castilleja applegatei +
variety +
green +  and yellow +
shallow;deep +
incurved +  and ascending +
reduced +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
1.1 cm11 mm <br />0.011 m <br /> (1.3 cm13 mm <br />0.013 m <br />) +