Castilleja applegatei var. pinetorum

(Fernald) N. H. Holmgren in A. Cronquist et al.

Intermount. Fl. 4: 486. 1984.

Common names: Wavy-leaved paintbrush
IllustratedEndemic
Basionyms: Castilleja pinetorum Fernald Erythea 6: 50. 1898
Synonyms: C. applegatei var. fragilis (Zeile) N. H. Holmgren C. applegatei subsp. pinetorum (Fernald) T. I. Chuang & Heckard C. wherryana Pennell
Treatment appears in FNA Volume 17. Treatment on page 589. Mentioned on page 583, 588, 594, 603, 648.

Herbs 1.2–6 dm. Stems unbranched, sometimes branched. Leaves linear to broadly lanceolate, 1–6 cm, 0–3 (–5) -lobed, apex narrowly acute to acuminate. Inflorescences 4–12 (–21 in fruit) × 1–4 cm; bracts distally red, red-orange, or scarlet, sometimes orange, pale orange, or yellow, rarely with a narrow yellowish band medially, 3 (–7) -lobed, central lobe apex obtuse, others obtuse to acute. Calyces proximally green, yellow, or whitish with green veins, lobe apices colored as bracts, sometimes with a yellow band between green and other color, 15–25 mm; abaxial and adaxial clefts 4–9 mm, 33–50% of calyx length, lateral 2.5–5 mm, 12–20% of calyx length; lobe apex acute to acuminate. Corollas 25–35 mm; tube 15–20 mm; beak 9–20 mm. 2n = 24, 48.


Phenology: Flowering (Apr–)May–Aug.
Habitat: Open forests, rocky slopes, canyons, sagebrush hillsides.
Elevation: 300–3600 m.

Distribution

Calif., Idaho, Nev., Oreg.

Discussion

Variety pinetorum is the most widespread and common representative of the Castilleja applegatei complex. It replaces var. breweri in the Sierra Nevada at low to moderate elevations. On the western flank of Steens Mountain, Harney County, Oregon, var. pinetorum occurs almost exclusively in a yellow-bracted form and has been confused with C. glandulifera. Variety pinetorum hybridizes with C. peckiana in Harney County, Oregon.

Selected References

None.

Lower Taxa

None.

0.25 cm2.5 mm <br />0.0025 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
green +  and whitish or yellow +
exserted +  and included +
protuberant +  and inconspicuous +
0.1 cm1 mm <br />0.001 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
J. Mark Egger +, Peter F. Zika +, Barbara L. Wilson +, Richard E. Brainerd +  and Nick Otting +
(Fernald) N. H. Holmgren in A. Cronquist et al. +
yellow +  and yellowish +
Castilleja pinetorum +
long-exserted +
0.9 cm9 mm <br />0.009 m <br /> (2 cm20 mm <br />0.02 m <br />) +
not +, leathery +  and fleshy +
yellow;pale orange;yellow;pale orange;orange;scarlet;red-orange;scarlet;red-orange;red +
3(-7)-lobed +
whitish +, yellow +  and green +
2-lobed +, lobed +  and tubular +
taprooted;fibrous-rooted or rhizomatous +
fleshy +  and woody +
Wavy-leaved paintbrush +
3.5 cm35 mm <br />0.035 m <br /> (4.1 cm41 mm <br />0.041 m <br />) +
purple +, pink +, red +, orange +, yellow +, white +, green +  and pale greenish +
curved +  and straight +
2.5 cm25 mm <br />0.025 m <br /> (3.5 cm35 mm <br />0.035 m <br />) +
Calif. +, Idaho +, Nev. +  and Oreg. +
300–3600 m. +
spreading-hairy +  and glabrous +
2 +  and 1 +
indehiscent +, septicidal +, loculicidal +  and dehiscence +
Open forests, rocky slopes, canyons, sagebrush hillsides. +
sparse +  and dense +
stipitate-glandular +
axillary +  and terminal +
spiral +, whorled +, alternate +  and opposite +
green +  and purplish or brown +
deciduous +
basal +  and cauline +
0-3(-5)-lobed;linear;broadly lanceolate +
1 cm10 mm <br />0.01 m <br /> (6 cm60 mm <br />0.06 m <br />) +
not fleshy +
1.5 cm15 mm <br />0.015 m <br /> (2.5 cm25 mm <br />0.025 m <br />) +
arising +, spreading +  and ascending +
3 +, 2 +  and 4 +
triangular +, oblong +, lanceolate-acuminate +, lanceolate +  and broadly lanceolate +
expanded +
(0-)3(-5)-lobed +  and wavy +
obtuse;acute +
tenuinucellate +  and unitegmic +
campylotropous-like +
Flowering (Apr–)May–Aug. +
Intermount. Fl. +
20 +  and 100 +
trapezoidal +, oblong-ovoid +, pyramidal +  and ovoid +
Illustrated +  and Endemic +
non-secund +  and secund +
elongate +
subequal +
branched +  and unbranched +
sprawling +  and prostrate +
aerial +  and subterranean +
decumbent +, ascending +  and erect +
not fleshy +
2-lobed +  and entire +
C. applegatei var. fragilis +, C. applegatei subsp. pinetorum +  and C. wherryana +
Castilleja applegatei var. pinetorum +
Castilleja applegatei +
variety +
green +  and yellow +
shallow;deep +
incurved +  and ascending +
reduced +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
1.5 cm15 mm <br />0.015 m <br /> (2 cm20 mm <br />0.02 m <br />) +