Castilleja arachnoidea

Greenman

Bot. Gaz. 53: 510. 1912.

Common names: Cobwebby or cotton paintbrush
IllustratedEndemic
Synonyms: Castilleja arachnoidea subsp. shastensis Pennell C. eastwoodiana Pennell C. filifolia Eastwood C. payneae Eastwood C. pumicicola Pennell
Treatment appears in FNA Volume 17. Treatment on page 590. Mentioned on page 568, 571, 583, 655.

Herbs, perennial, 0.6–2 (–3) dm; from branched, sometimes elongate, creeping basal stems; with a taproot. Stems solitary or few-to-many, decumbent-ascending, unbranched, sometimes branched, hairs dense, reflexed-spreading, matted, long, soft, unbranched, eglandular, white-woolly. Leaves green to purple or steel gray, lanceolate-linear or narrowly lanceolate, (1–) 2–4 (–6) cm, not fleshy, margins plane, involute, 3 (–5) -lobed, apex acute; lobes spreading, linear to lanceolate, apex acute to rounded. Inflorescences erect to ascending, 3–12 × 1–2.5 cm; bracts proximally greenish to dull reddish-brown, rarely dull red or dull orange, distally dull red, deep rusty red, or yellow to pale-yellow, sometimes deep pink, dull rose, pale salmon, or dull light orange, rarely green, dull reddish, or dull orange throughout, aging browner and/or more orange, lanceolate or elliptic-lanceolate to obovate, (0–) 3–5-lobed; lobes usually ascending, linear, lanceolate, oblanceolate, or spatulate, long, arising near or below mid length, central lobe apex rounded to truncate, others obtuse to rounded. Calyces colored as bracts, proximal part sometimes paler, (10–) 12–19 mm; abaxial and adaxial clefts 4–8 (–10) mm, 33–50% of calyx length, all 4 clefts subequal; lobes linear to lanceolate, apex acute. Corollas straight or ± curved, 12–20 mm; tube 9–15 mm; beak subequal to calyx or slightly exserted, adaxially green or yellow-green, (2–) 3–5 mm, ± densely puberulent, hairs often crisped; abaxial lip yellow, greenish, pink, or red-violet, moderately conspicuous, exserted to barely included in calyx; pouches shallow, prominent, 2–4 mm, ca. 100% as long as beak; teeth erect, red, pink, or pale-yellow, 1 mm. 2n = 24.


Phenology: Flowering (May–)Jun–Aug(–Sep).
Habitat: Pumice flats, sandy, gravelly, or rocky slopes, ridges and open summits.
Elevation: 1300–3300 m.

Discussion

Castilleja arachnoidea is a remarkably variable species in form and color, but inflorescence colors are often uniform within a population. It is sometimes confused with C. schizotricha, but the two differ in stature, lobing of the leaves and bracts, bract coloration, and inflation of the abaxial lip. They are easily separated by the simple hairs and pale yellow to brick red inflorescences of C. arachnoidea, compared to the pink-purple to deep purple inflorescences and branched hairs of C. schizotricha.

The names Castilleja eastwoodiana and C. filifolia represent yellow color forms of C. arachnoidea. A race on the pumice plains of Mt. Shasta in the Cascade Range in California is distinctive.

Selected References

None.

Lower Taxa

None.

"arising" is not a number.

... more about "Castilleja arachnoidea"
0.8 cm8 mm <br />0.008 m <br /> (1 cm10 mm <br />0.01 m <br />) +
0.4 cm4 mm <br />0.004 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
red-violet +, pink +, greenish +  and yellow +
exserted +  and barely included +
Acute (?) +  and Rounded (?) +
J. Mark Egger +, Peter F. Zika +, Barbara L. Wilson +, Richard E. Brainerd +  and Nick Otting +
Greenman +
subequal +
not +, leathery +  and fleshy +
orange;browner;orange;dull;reddish;dull;grorange;browner;orange;dull;reddish;dull;green;light orange;dull;pale salmon;orange;dull;reddish;dull;green;light orange;dull;pale salmon;orange;dull;reddish;dull;green;light orange;dull;pale salmon;orange;dull;reddish;dull;green;light orange;dull;pale salmon;orange;dull;reddish;dull;green;light orange;dull;pale salmon;rose;dull;pink;deep;proximally greenish;dull reddish-brown rarely dull red or dull orange distally dull red deep rusty red or yellowdistally dull red deep rusty red or yellow +
(0-)3-5-lobed +, elliptic-lanceolate +  and obovate +
2-lobed +, lobed +  and tubular +
taprooted;fibrous-rooted or rhizomatous +
fleshy +  and woody +
subequal +
Cobwebby or cotton paintbrush +
purple +, pink +, red +, orange +, yellow +, white +, green +  and pale greenish +
curved +  and straight +
1.2 cm12 mm <br />0.012 m <br /> (2 cm20 mm <br />0.02 m <br />) +
Calif. +, Nev. +  and Oreg. +
1300–3300 m. +
spreading-hairy +  and glabrous +
2 +  and 1 +
indehiscent +, septicidal +, loculicidal +  and dehiscence +
Pumice flats, sandy, gravelly, or rocky slopes, ridges and open summits. +
eglandular +  and unbranched +
0.2 cm2 mm <br />0.002 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
yellow-green +  and green +
reflexed-spreading +
puberulent +  and white-woolly +
crisped +
0.3 cm3 mm <br />0.003 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
axillary +  and terminal +
spiral +, whorled +, alternate +  and opposite +
4 cm40 mm <br />0.04 m <br /> (6 cm60 mm <br />0.06 m <br />) +
gray +, steel +, green +  and purple +
deciduous +
basal +  and cauline +
lanceolate;lanceolate-linear +
2 cm20 mm <br />0.02 m <br /> (4 cm40 mm <br />0.04 m <br />) +
not fleshy +
ascending +  and spreading +
3 +, 2 +  and 4 +
linear +  and lanceolate +
subequal +
3(-5)-lobed +  and plane +
obtuse;rounded +
tenuinucellate +  and unitegmic +
campylotropous-like +
Flowering (May–)Jun–Aug(–Sep). +
shallow +
0.2 cm2 mm <br />0.002 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
1 cm10 mm <br />0.01 m <br /> (1.2 cm12 mm <br />0.012 m <br />) +
20 +  and 100 +
trapezoidal +, oblong-ovoid +, pyramidal +  and ovoid +
Illustrated +  and Endemic +
non-secund +  and secund +
elongate +
subequal +
branched +  and unbranched +
sprawling +  and prostrate +
aerial +  and subterranean +
decumbent-ascending +
not fleshy +
2-lobed +  and entire +
Castilleja arachnoidea subsp. shastensis +, C. eastwoodiana +, C. filifolia +, C. payneae +  and C. pumicicola +
Castilleja arachnoidea +
Castilleja +
species +
pale-yellow +, pink +  and red +
shallow;deep +
reduced +
1 cm10 mm <br />0.01 m <br /> (?) +
0.9 cm9 mm <br />0.009 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +