Castilleja cusickii

Greenman

Bot. Gaz. 25: 267. 1898.

IllustratedEndemic
Synonyms: Castilleja lutea A. Heller
Treatment appears in FNA Volume 17. Treatment on page 600. Mentioned on page 574, 601, 611.

Herbs, perennial, (1–) 1.5–5 (–6) dm; from a small, woody caudex; with a taproot or stout, branched roots. Stems solitary or few to several, erect, often decumbent at base, unbranched, sometimes branched, hairs spreading, long, soft, eglandular, mixed with shorter, sometimes stiff, stipitate-glandular ones. Leaves green, sometimes with prominent red-purple veins, lanceolate-linear to broadly lanceolate, 2.5–4.5 (–7) cm, not fleshy, margins plane, flat, (0–) 3–5 (–9) -lobed, apex acuminate to acute; lobes spreading-ascending, narrowly lanceolate, arising at or above mid length, apex acute. Inflorescences 3.5–26 × 1.5–3.5 cm; bracts pale green, pale greenish yellow, or pale-yellow throughout, or proximally pale green, pale greenish yellow, pale-yellow, or reddish purple, distally white, yellow, pink, dull purple, or dull reddish purple, sometimes with a purple band below that, sometimes with veins darker than background color, lanceolate to oblong, 0–5 (–9) -lobed; lobes ascending or spreading, linear to narrowly lanceolate, often short, arising above mid length, central lobe apex rounded, lateral ones acute to rounded. Calyces green, pale green, or pale-yellow, lobes yellow to pale-yellow, 20–30 mm; abaxial and adaxial clefts 6.6–14 mm, 33–50% of calyx length, deeper than laterals, lateral 0–4 (–4.5) mm, 0–15% of calyx length; lobes broadly rounded to triangular, apex obtuse to rounded. Corollas straight, 19–25 (–28) mm; tube (13–) 15–20 mm; abaxial lip included, beak not or slightly exserted; beak adaxially green to yellow-green, 4–7 mm; abaxial lip yellow, prominent, inflated, 2–4.5 mm, ca. 75% as long as beak; teeth incurved to erect, white, 1.3–2.1 mm. 2n = 24.


Phenology: Flowering Apr–Aug.
Habitat: Moist meadows, swales, grasslands in sagebrush steppes, occasionally to subalpine.
Elevation: 500–2500 m.

Distribution

Alta., B.C., Idaho, Mont., Nev., Oreg., Wash., Wyo.

Discussion

Castilleja cusickii includes two morphologically distinct variants. One variant has a wider inflorescence, broader bracts, and wider bract lobes. These bracts often have highly contrasting purple veins, and some populations also have a purplish wash on the bracts. These plants have a range on the western edge of typical C. cusickii, though overlapping with more typical forms in western Idaho and adjacent northeastern Oregon and southeastern Washington. The name C. lutea is available for these variants, and both consistent morphological differences and preliminary results from ongoing genetic research on the plants (D. C. Tank, pers. comm.) indicate its resurrection may be justified. A second variant is distinguished by a pale pink-purple wash on the bracts and a narrow distribution in the meadows of the Bear River Valley, bordering northwestern Utah and adjacent southeastern Idaho. Only bract coloration appears to distinguish this form from typical C. cusickii, and the distinctive bract coloration is only more or less consistent in these populations.

Typical Castilleja cusickii appears to hybridize with C. gracillima in the Logan Valley, Grant County, Oregon. It is also reported by M. Ownbey (1959) to hybridize with C. miniata and C. rhexiifolia.

Selected References

None.

Lower Taxa

None.

"arising" is not a number.

... more about "Castilleja cusickii"
0.4 cm4 mm <br />0.004 m <br /> (0.45 cm4.5 mm <br />0.0045 m <br />) +
0 cm0 mm <br />0 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
0.2 cm2 mm <br />0.002 m <br /> (0.45 cm4.5 mm <br />0.0045 m <br />) +
Obtuse (?) +  and Rounded (?) +
J. Mark Egger +, Peter F. Zika +, Barbara L. Wilson +, Richard E. Brainerd +  and Nick Otting +
Greenman +
0.4 cm4 mm <br />0.004 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
not +, leathery +  and fleshy +
pink;yellow;white;reddish purple;pale-yellow;reddish purple;pale-yellow;yellow;pale greenish;pale green;pale-yellow;yellow;pale-yellow;yellow;pale greenish;pale green +
reddish purple +, dull +  and purple +
0-5(-9)-lobed +, lanceolate +  and oblong +
pale-yellow +, pale green +  and green +
2-lobed +, lobed +  and tubular +
taprooted;fibrous-rooted or rhizomatous +
fleshy +  and woody +
2.5 cm25 mm <br />0.025 m <br /> (2.8 cm28 mm <br />0.028 m <br />) +
purple +, pink +, red +, orange +, yellow +, white +, green +  and pale greenish +
1.9 cm19 mm <br />0.019 m <br /> (2.5 cm25 mm <br />0.025 m <br />) +
Alta. +, B.C. +, Idaho +, Mont. +, Nev. +, Oreg. +, Wash. +  and Wyo. +
500–2500 m. +
spreading-hairy +  and glabrous +
2 +  and 1 +
indehiscent +, septicidal +, loculicidal +  and dehiscence +
Moist meadows, swales, grasslands in sagebrush steppes, occasionally to subalpine. +
stipitate-glandular +
axillary +  and terminal +
spiral +, whorled +, alternate +  and opposite +
4.5 cm45 mm <br />0.045 m <br /> (7 cm70 mm <br />0.07 m <br />) +
deciduous +
basal +  and cauline +
lanceolate-linear;broadly lanceolate +
2.5 cm25 mm <br />0.025 m <br /> (4.5 cm45 mm <br />0.045 m <br />) +
not fleshy +
yellow +  and pale-yellow +
spreading +, ascending +  and spreading-ascending +
3 +, 2 +  and 4 +
broadly rounded +  and triangular +
subequal +
(0-)3-5(-9)-lobed +  and plane +
stipitate-glandular +
tenuinucellate +  and unitegmic +
campylotropous-like +
Flowering Apr–Aug. +
20 +  and 100 +
trapezoidal +, oblong-ovoid +, pyramidal +  and ovoid +
Illustrated +  and Endemic +
non-secund +  and secund +
elongate +
subequal +
branched +  and unbranched +
aerial +  and subterranean +
not fleshy +
2-lobed +  and entire +
Castilleja lutea +
Castilleja cusickii +
Castilleja +
species +
shallow;deep +
incurved +  and erect +
reduced +
0.13 cm1.3 mm <br />0.0013 m <br /> (0.21 cm2.1 mm <br />0.0021 m <br />) +
1.3 cm13 mm <br />0.013 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
1.5 cm15 mm <br />0.015 m <br /> (2 cm20 mm <br />0.02 m <br />) +
darker than background color +