Castilleja exserta var. latifolia

(S. Watson) J. M. Egger

Phytologia 90: 70. 2008.

Endemic
Basionyms: Orthocarpus purpurascens var. latifolius S. Watson Botany (Fortieth Parallel), 458. 1871
Synonyms: Castilleja exserta subsp. latifolia (S. Watson) T. I. Chuang & Heckard
Treatment appears in FNA Volume 17. Treatment on page 607. Mentioned on page 606.

Herbs 0.6–3.3 dm. Inflorescences 1.5–12 cm; bracts: distal apices pink to pale-purple, sometimes white, 5–7 mm wide, 3–5 (–7) -lobed; lobes linear to narrowly spatulate, less than 2 × 1–2 mm. Calyces 17.5–23 mm; abaxial clefts 4–6 mm, adaxial 9–16 mm, abaxial 40–50% of calyx length, adaxial 50–70% of calyx length, lateral 2.5–3 mm, 15% of calyx length. Corollas 20–27 mm; tube 15–20 mm; beak 5–7 mm; abaxial lip pink proximally, distal 1/3 white to yellow, becoming deep pink after anthesis, 3–5 mm.


Phenology: Flowering (Mar–)Apr–Aug.
Habitat: Coastal grasslands, dunes, sandy flats, headlands, sandy to rocky slopes.
Elevation: 0–500 m.

Discussion

Variety latifolia is a close associate of Artemisia pycnocephala along the coast of central and northern California. Sand dune populations in the vicinity of Humboldt Bay and occasionally elsewhere are unique in having almost uniformly greenish bracts, usually with white apices, and a less-inflated abaxial lip on the corollas. The inflorescences of this form also become anthcyanic very rapidly, with much of the coloration fading quickly to deep, dull reddish brown, even while flowering continues. These distinctive plants are the subject of ongoing research. The northernmost population of this form of var. latifolia occurs north of Crescent City, in northern Del Norte County, and this variety should be looked for in adjacent southwestern Oregon.

Selected References

None.

Lower Taxa

None.

"rounded" is not a number."adaxial" is not a number. "length" is not a number.

0.4 cm4 mm <br />0.004 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
pink +, white +  and yellow +
0.3 cm3 mm <br />0.003 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
0.25 cm2.5 mm <br />0.0025 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
Rounded (?) +  and Acute (?) +
J. Mark Egger +, Peter F. Zika +, Barbara L. Wilson +, Richard E. Brainerd +  and Nick Otting +
(S. Watson) J. M. Egger +
Orthocarpus purpurascens var. latifolius +
0.5 cm5 mm <br />0.005 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
not +, leathery +  and fleshy +
white;light purple;magenta;lavender;pink;wwhite;light purple;magenta;lavender;pink;white;brownish purple;white;light purple;magenta;lavender;pink;white;brownish purple;white;light purple;magenta;lavender;pink;white;brownish purple;white;light purple;magenta;lavender;pink;white;brownish purple;white;light purple;magenta;lavender;pink;white;brownish purple;white;light purple;magenta;lavender;pink;white;brownish purple;white;light purple;magenta;lavender;pink;white;brownish purple;dark purple;greenishwhite;brownish purple;dark purple;greenish +
(3-)5(-9)-lobed +, lanceolate +  and elliptic or narrowly ovate +
2-lobed +, lobed +  and tubular +
1.75 cm17.5 mm <br />0.0175 m <br /> (2.3 cm23 mm <br />0.023 m <br />) +
taprooted;fibrous-rooted or rhizomatous +
fleshy +  and woody +
adaxial +  and length +
purple +, pink +, red +, orange +, yellow +, white +, green +  and pale greenish +
2 cm20 mm <br />0.02 m <br /> (2.7 cm27 mm <br />0.027 m <br />) +
3-5(-7)-lobed +
0.5 cm5 mm <br />0.005 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
aging +  and purple spots +
0–500 m. +
spreading-hairy +  and glabrous +
2 +  and 1 +
indehiscent +, septicidal +, loculicidal +  and dehiscence +
Coastal grasslands, dunes, sandy flats, headlands, sandy to rocky slopes. +
much-branched +, unbranched +  and eglandular +
stipitate-glandular +
axillary +  and terminal +
1.5 cm15 mm <br />0.015 m <br /> (12 cm120 mm <br />0.12 m <br />) +
spiral +, whorled +, alternate +  and opposite +
5 cm50 mm <br />0.05 m <br /> (7.7 cm77 mm <br />0.077 m <br />) +
purple +  and brownish +
deciduous +
basal +  and cauline +
ovate;orbicular +
1 cm10 mm <br />0.01 m <br /> (5 cm50 mm <br />0.05 m <br />) +
not fleshy +
0 cm0 mm <br />0 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
above mid lobes +, arising +, ascending +  and spreading +
3 +, 2 +  and 4 +
linear +  and narrowly spatulate +
subequal +
1mm;2mm +
villous-hairy +
(0-)3-9(-11)-lobed +  and plane +
short-stipitate-glandular +
tenuinucellate +  and unitegmic +
campylotropous-like +
Flowering (Mar–)Apr–Aug. +
0.3 cm3 mm <br />0.003 m <br /> (0.9 cm9 mm <br />0.009 m <br />) +
0.3 cm3 mm <br />0.003 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
20 +  and 100 +
trapezoidal +, oblong-ovoid +, pyramidal +  and ovoid +
non-secund +  and secund +
elongate +
subequal +
branched +  and unbranched +
sprawling +  and prostrate +
aerial +  and subterranean +
erect +  and ascending +
not fleshy +
2-lobed +  and entire +
Castilleja exserta subsp. latifolia +
Castilleja exserta var. latifolia +
Castilleja exserta +
variety +
yellow +  and white +
shallow;deep +
reduced +
1.5 cm15 mm <br />0.015 m <br /> (2 cm20 mm <br />0.02 m <br />) +