Castilleja fraterna

Greenman

Bot. Gaz. 48: 147. 1909.

Common names: Fraternal paintbrush
EndemicConservation concern
Treatment appears in FNA Volume 17. Treatment on page 609. Mentioned on page 580, 597.

Herbs, perennial, 0.8–2.5 dm; from a woody caudex; with a taproot. Stems few-to-many, ± curved at base, ascending or erect distally, unbranched, hairs moderately dense, spreading, medium length and long, soft, mixed with shorter stipitate-glandular ones. Leaves green, sometimes purple-tinged, lanceolate, broadly lanceolate, or narrowly ovate, 1–4.3 cm, not fleshy, margins plane, sometimes ± wavy, ± involute, 0–3-lobed, apex acute to acuminate; lobes ascending, often narrowly oblanceolate, apex acute to acuminate. Inflorescences 3–8.5 × 1–4 cm; bracts greenish or bright to sometimes dull red, sometimes scarlet, orange, or pale-pink throughout, or proximally greenish, distally as above, ovate to elliptic, (0–) 3-lobed; lobes ascending, lanceolate to narrowly triangular, short, arising above mid length, apex acute to obtuse or acuminate. Calyces brightly, conspicuously colored for at least distal 2/3, often throughout, providing much of inflorescence coloration, proximally pale whitish to pale-pink, distally colored as in distal portion of bracts, sometimes colored throughout as in distal portion of bracts, 15–30 mm; abaxial and adaxial clefts 7–14 mm, 33–50% of calyx length, deeper than laterals, lateral 1–5 mm, 5–15% of calyx length; lobes narrowly to broadly triangular, apex rounded to acute. Corollas straight or slightly curved, 20–40 mm; tube 27 mm; abaxial lip often visible in abaxial cleft, sometimes exserted, beak usually exserted from calyx; beak adaxially green to yellow, 8–14 mm; abaxial lip red, black, green, white, or green-and-white, ± prominent, slightly rounded, 1.5–5 mm, ca. 33% as long as beak; teeth ascending, green, bright red, pink, white, or yellow, 0.5–3 mm.


Phenology: Flowering Jul–Aug(–Sep).
Habitat: Moist or dry rocky slopes and flats, ridges, talus, dwarf willow mats, subalpine to alpine, rarely along stream channels at lower elevations, over sedimentary rocks, often limy.
Elevation: 2000–2900 m.

Discussion

Castilleja fraterna is endemic to one ridge system in the Wallowa Mountains of northeastern Oregon. A parallel ridge system in the range has a second Wallowa Mountains endemic, C. rubida. Castilleja fraterna is colored similarly to C. miniata, but the petaloid teeth of the abaxial corolla lip are more like C. chrysantha. It is possible that C. fraterna was derived through hybridization, though its chromosome number is unknown. It resembles the newly described C. kerryana in Montana.

Selected References

None.

Lower Taxa

None.

"arising" is not a number.

... more about "Castilleja fraterna"
green-and-white +, white +, green +, black +  and red +
prominent +  and visible +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
Rounded (?) +  and Acute (?) +
J. Mark Egger +, Peter F. Zika +, Barbara L. Wilson +, Richard E. Brainerd +  and Nick Otting +
Greenman +
0.8 cm8 mm <br />0.008 m <br /> (1.4 cm14 mm <br />0.014 m <br />) +
not +, leathery +  and fleshy +
greenish;pale-pink;orange;pale-pink;orange;scarlet;to bright sometimes sometimes dull red;greenish +
or +  and sometimes dull +
(0-)3-lobed +, ovate +  and elliptic +
coloration +
lateral +  and deeper +
0.1 cm1 mm <br />0.001 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
colored +, proximally pale whitish +  and pale-pink +
2-lobed +, lobed +  and tubular +
1.5 cm15 mm <br />0.015 m <br /> (3 cm30 mm <br />0.03 m <br />) +
taprooted;fibrous-rooted or rhizomatous +
fleshy +  and woody +
Fraternal paintbrush +
purple +, pink +, red +, orange +, yellow +, white +, green +  and pale greenish +
curved +  and straight +
2 cm20 mm <br />0.02 m <br /> (4 cm40 mm <br />0.04 m <br />) +
2000–2900 m. +
spreading-hairy +  and glabrous +
2 +  and 1 +
indehiscent +, septicidal +, loculicidal +  and dehiscence +
Moist or dry rocky slopes and flats, ridges, talus, dwarf willow mats, subalpine to alpine, rarely along stream channels at lower elevations, over sedimentary rocks, often limy. +
much-branched +, unbranched +  and eglandular +
stipitate-glandular +
axillary +  and terminal +
spiral +, whorled +, alternate +  and opposite +
purple-tinged +  and green +
deciduous +
basal +  and cauline +
ovate;lanceolate;ovate;lanceolate;lanceolate +
1 cm10 mm <br />0.01 m <br /> (4.3 cm43 mm <br />0.043 m <br />) +
not fleshy +
3 +, 2 +  and 4 +
triangular +, lanceolate +  and narrowly triangular +
subequal +
0-3-lobed +, wavy +  and plane +
tenuinucellate +  and unitegmic +
campylotropous-like +
Flowering Jul–Aug(–Sep). +
20 +  and 100 +
trapezoidal +, oblong-ovoid +, pyramidal +  and ovoid +
Endemic +  and Conservation concern +
non-secund +  and secund +
elongate +
subequal +
sprawling +  and prostrate +
aerial +  and subterranean +
erect +  and ascending +
not fleshy +
2-lobed +  and entire +
Euchroma +  and Oncorhynchus +
Castilleja fraterna +
Castilleja +
species +
yellow +, white +, pink +, bright red +  and green +
shallow;deep +
reduced +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
27 cm270 mm <br />0.27 m <br /> (?) +