Castilleja grisea

Dunkle

Bull. S. Calif. Acad. Sci. 42: 31. 1943.

Common names: San Clemente Island paintbrush
EndemicConservation concern
Synonyms: Castilleja hololeuca subsp. grisea (Dunkle) Munz
Treatment appears in FNA Volume 17. Treatment on page 611. Mentioned on page 567, 570, 572, 612.

Subshrubs or shrubs, perennial, 4–6 (–10) dm, to 20 dm wide; from a woody caudex; with thick, woody roots. Stems many, spreading and ascending, much-branched, with short, leafy axillary shoots, proximal stems 1+ cm wide, hairs dense, spreading, matted, white, medium length, soft, stellate, sometimes mixed with short-stipitate-glandular ones especially near inflorescence, partially obscuring surface. Leaves ash gray, linear to narrowly lanceolate, 1–6.5 cm, not fleshy, margins plane or slightly wavy, involute, 0–7-lobed, apex rounded to obtuse, surfaces almost obscured by hairs, at least when young; lobes spreading to ascending, linear to narrowly oblong, apex acute to acuminate. Inflorescences (3–) 7.5–11.5 × 1.5–3.5 cm; bracts proximally greenish, distally ± pale-yellow, proximals sometimes greenish throughout, linear to narrowly lanceolate or narrowly oblong, (3–) 5–7-lobed; lobes spreading, linear to narrowly oblong, long, proximal lobes arising below mid length, apex obtuse to acute. Calyces colored as bracts, 10–20 mm; abaxial and adaxial clefts 5.5–7 mm, 50% of calyx length, deeper than laterals, lateral 0–1 mm, 0–8% of calyx length; lobes oblong, apex truncate to rounded or obtuse, rarely emarginate, inner surface glandular. Corollas curved in proximal 1/3, 15–25 mm; tube 11 mm; abaxial lip not exserted, sometimes visible in abaxial cleft, beak exserted; beak adaxially green or dull yellow, 7–9 mm; abaxial lip deep green, reduced, 1–2 mm, less than 25% as long as beak; teeth incurved, green, 0.7–1 mm. 2n = 24.


Phenology: Flowering Feb–Jun.
Habitat: Coastal terraces and slopes, cliffs and canyon walls, coastal sage scrub.
Elevation: 0–500 m.

Discussion

Castilleja grisea is endemic to a small portion of San Clemente Island in Los Angeles County, which is managed by the U.S. Navy. It was one of the first plants listed as endangered under the Endangered Species Act of the United States, and its near extinction was due to grazing by feral goats. Plants responded well when the goats were removed, spreading from their last refuge on inaccessible cliffs onto gentle terrain at the southern end of the island, near the type locality, and are now locally fairly common. Ungrazed old shrubs can reach 1 m in height and 2 m in breadth and often have a thick woody trunk. The stellate pubescence is unusual in Castilleja.

Castilleja grisea is in the Center for Plant Conservation’s National Collection of Endangered Plants.

Selected References

None.

Lower Taxa

None.

... more about "Castilleja grisea"
0.1 cm1 mm <br />0.001 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
J. Mark Egger +, Peter F. Zika +, Barbara L. Wilson +, Richard E. Brainerd +  and Nick Otting +
Dunkle +
Yellow (?) +, Dull (?) +  and Green (?) +
0.7 cm7 mm <br />0.007 m <br /> (0.9 cm9 mm <br />0.009 m <br />) +
not +, leathery +  and fleshy +
pale-yellow;greenish +
lateral +  and deeper +
0 cm0 mm <br />0 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
2-lobed +, lobed +  and tubular +
1 cm10 mm <br />0.01 m <br /> (2 cm20 mm <br />0.02 m <br />) +
taprooted;fibrous-rooted or rhizomatous +
fleshy +  and woody +
San Clemente Island paintbrush +
purple +, pink +, red +, orange +, yellow +, white +, green +  and pale greenish +
1.5 cm15 mm <br />0.015 m <br /> (2.5 cm25 mm <br />0.025 m <br />) +
0–500 m. +
spreading-hairy +  and glabrous +
2 +  and 1 +
indehiscent +, septicidal +, loculicidal +  and dehiscence +
Coastal terraces and slopes, cliffs and canyon walls, coastal sage scrub. +
much-branched +, unbranched +  and eglandular +
stipitate-glandular +
3 cm30 mm <br />0.03 m <br /> (7.5 cm75 mm <br />0.075 m <br />) +
axillary +  and terminal +
spiral +, whorled +, alternate +  and opposite +
deciduous +
basal +  and cauline +
linear;narrowly lanceolate +
1 cm10 mm <br />0.01 m <br /> (6.5 cm65 mm <br />0.065 m <br />) +
not fleshy +
spreading +  and ascending +
3 +, 2 +  and 4 +
oblong +, linear +  and narrowly oblong +
subequal +
0-7-lobed +, wavy +  and plane +
short-stipitate-glandular +
tenuinucellate +  and unitegmic +
campylotropous-like +
Flowering Feb–Jun. +
(3-)5-7-lobed;linear;narrowly lanceolate or narrowly oblong +
Bull. S. Calif. Acad. Sci. +
20 +  and 100 +
trapezoidal +, oblong-ovoid +, pyramidal +  and ovoid +
Endemic +  and Conservation concern +
non-secund +  and secund +
elongate +
subequal +
much-branched +
sprawling +  and prostrate +
aerial +  and subterranean +
ascending +  and spreading +
not fleshy +
2-lobed +  and entire +
Castilleja hololeuca subsp. grisea +
Castilleja grisea +
Castilleja +
species +
shallow;deep +
reduced +
0.07 cm0.7 mm <br />7.0e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
11 cm110 mm <br />0.11 m <br /> (?) +
shrub +  and subshrub +