IllustratedEndemic
Synonyms: Castilleja hyetophila Pennell C. inconstans Standley
Treatment appears in FNA Volume 17. Treatment on page 628. Mentioned on page 588, 605, 627, 629, 630.

Herbs 1.6–6.5 dm; with a taproot or with slender, branched roots from a rhizome. Stems glabrous or sparsely hairy proximally and medially, subglabrous or sparsely hairy distally, hairs spreading, short to long, soft, eglandular, rarely stipitate-glandular. Leaves spreading to ascending, linear to lanceolate, narrowly elliptic, or narrowly oblong, thin and not fleshy, margins plane, rarely wavy, flat to slightly involute, leaf sometimes recurved downward, 0 (–3) -lobed, apex acute to acuminate, sometimes obtuse. Bracts distally scarlet, red, orange-red, or pale orange, sometimes pink, pink-purple, magenta, pale orange, yellow, greenish yellow, or white, 0–3 (–5) -lobed, central lobe with 3–5 small teeth at apex; central lobe apex obtuse, rounded to truncate, lateral ones acute to rounded. Calyces 15–36 mm; abaxial and adaxial clefts 8–20 mm, ca. 40% of calyx length, subequal or abaxial longer, lateral 1–9 mm, 15–20% of calyx length; lobes narrowly lanceolate to narrowly triangular or linear, apex acute, sometimes acuminate or obtuse. Corollas 20–48 mm; beak 9–25 mm; abaxial lip deep green or deep purple, 0.5–3.5 mm, 10–15% as long as beak, to ca. 33% as long as beak in some populations. 2n = 24, 48, 72, 96, 120.


Phenology: Flowering Mar–Sep(–Oct).
Habitat: Moist or wet meadows, stream banks, shores, open forests, rocky slopes, roadsides, forest edges.
Elevation: 0–3700 m.

Distribution

Alta., B.C., Man., Ont., Sask., Alaska, Ariz., Calif., Colo., Idaho, Mont., Nev., N.Mex., Oreg., Utah, Wash., Wyo.

Discussion

Variety miniata is much more widespread than the other varieties of Castilleja miniata. Occasional hybrids are encountered between this variety and a wide range of species. Apparent hybrid swarms with C. rhexiifolia and/or C. septentrionalis are occasional in the Rocky Mountains. In the Pacific Northwest and across the prairie provinces of Canada, populations with relatively short inflorescences and pink bracts deserve further study. Likewise, populations of pink to pink-purple plants with shorter corollas and corolla beaks are found on boggy ground in the Peace River region of northwestern Alberta and adjacent British Columbia. A slightly rhizomatous form from the coast of British Columbia to southeastern Alaska was named C. hyetophila. The precise relationship of the rhizomatous form with var. dixonii and with C. miniata as a whole is still unclear and merits comprehensive study.

Selected References

None.

Lower Taxa

None.

"thin" is not a number."medium" is not a number.

120 +, 96 +, 72 +, 48 +  and 24 +
0.8 cm8 mm <br />0.008 m <br /> (1.2 cm12 mm <br />0.012 m <br />) +
0.3 cm3 mm <br />0.003 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
purple +, deep +  and green +
ascending +  and incurved +
not inflated +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.35 cm3.5 mm <br />0.0035 m <br />) +
Obtuse (?) +, Acuminate (?) +  and Acute (?) +
J. Mark Egger +, Peter F. Zika +, Barbara L. Wilson +, Richard E. Brainerd +  and Nick Otting +
Douglas ex Hooker +
0.9 cm9 mm <br />0.009 m <br /> (1.4 cm14 mm <br />0.014 m <br />) +
0.9 cm9 mm <br />0.009 m <br /> (2.5 cm25 mm <br />0.025 m <br />) +
not +, leathery +  and fleshy +
white;greenish yellow;white;greenish yellow;yellow;pale orange;magenta;pink-purple;pink;pale orange;orange-red;pale orange;orange-red;red;scarlet +
0-3(-5)-lobed +
0.1 cm1 mm <br />0.001 m <br /> (0.9 cm9 mm <br />0.009 m <br />) +
2-lobed +, lobed +  and tubular +
subequal +
1.5 cm15 mm <br />0.015 m <br /> (3.6 cm36 mm <br />0.036 m <br />) +
taprooted;fibrous-rooted or rhizomatous +
fleshy +  and woody +
0-5(-7)-lobed +, lanceolate +  and oblong-ovate +
purple +, pink +, red +, orange +, yellow +, white +, green +  and pale greenish +
2 cm20 mm <br />0.02 m <br /> (4.8 cm48 mm <br />0.048 m <br />) +
Alta. +, B.C. +, Man. +, Ont. +, Sask. +, Alaska +, Ariz. +, Calif. +, Colo. +, Idaho +, Mont. +, Nev. +, N.Mex. +, Oreg. +, Utah +, Wash. +  and Wyo. +
0–3700 m. +
spreading-hairy +  and glabrous +
2 +  and 1 +
indehiscent +, septicidal +, loculicidal +  and dehiscence +
Moist or wet meadows, stream banks, shores, open forests, rocky slopes, roadsides, forest edges. +
stipitate-glandular +
short to long +
15 cm150 mm <br />0.15 m <br /> (22 cm220 mm <br />0.22 m <br />) +
axillary +  and terminal +
spiral +, whorled +, alternate +  and opposite +
8 cm80 mm <br />0.08 m <br /> (9.5 cm95 mm <br />0.095 m <br />) +
green +  and purple +
deciduous +
basal +  and cauline +
0(-3)-lobed;linear;lanceolate narrowly elliptic or narrowly oblong +
3 cm30 mm <br />0.03 m <br /> (8 cm80 mm <br />0.08 m <br />) +
not fleshy +
above mid lobes +, arising +, erect +  and ascending-spreading +
3 +, 2 +  and 4 +
narrowly lanceolate +  and narrowly triangular or linear +
subequal +
flat +  and slightly involute +
tenuinucellate +  and unitegmic +
campylotropous-like +
0 cm0 mm <br />0 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
Flowering Mar–Sep(–Oct). +
Fl. Bor.-Amer. +
slender +
20 +  and 100 +
trapezoidal +, oblong-ovoid +, pyramidal +  and ovoid +
Illustrated +  and Endemic +
non-secund +  and secund +
elongate +
subequal +
branched +  and rooting +
creeping +  and decumbent +
aerial +  and subterranean +
erect +  and ascending +
not fleshy +
2-lobed +  and entire +
Castilleja hyetophila +  and C. inconstans +
Castilleja miniata var. miniata +
Castilleja miniata +
variety +
white +  and green +
shallow;deep +
erect +  and incurved +
3 +  and 5 +
0.07 cm0.7 mm <br />7.0e-4 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
1.2 cm12 mm <br />0.012 m <br /> (2.6 cm26 mm <br />0.026 m <br />) +