Castilleja minor var. spiralis

(Jepson) J. M. Egger

Phytologia 90: 74. 2008.

IllustratedEndemic
Basionyms: Castilleja spiralis Jepson Fl. W. Calif., 412. 1901
Synonyms: C. minor subsp. spiralis (Jepson) T. I. Chuang & Heckard C. stenantha subsp. spiralis (Jepson) Munz
Treatment appears in FNA Volume 17. Treatment on page 631. Mentioned on page 627, 630.

Herbs 5–7 dm. Stems: hairs dense, short, soft, mixed with shorter stipitate-glandular ones. Leaves linear or narrowly lanceolate, soft, margins plane or wavy. Bracts distally red, proximal narrowly lanceolate or oblanceolate, distal oblong to narrowly spatulate, apex rounded or obtuse, rarely acute. Calyces 15–27 mm; abaxial and adaxial clefts 7–9 mm, 35–50% of calyx length, lateral 0.5–2.5 mm, 5–25% of calyx length; lobes linear to narrowly lanceolate, apex acute to acuminate. Corollas 20–35 mm; beak and often abaxial lip exserted; beak adaxially yellowish or reddish-brown, 8–15 mm; abaxial lip red.


Phenology: Flowering Jun–Aug.
Habitat: Stream banks, damp flats and seeps, over serpentine.
Elevation: 0–3000 m.

Discussion

Variety spiralis is limited to a few highly serpentine sites in central California. Its range slightly overlaps var. stenantha, but within populations there is little or no intergradation in the distinguishing characters. Variety spiralis is separable from the other varieties of Castilleja minor by a combination of obtuse to rounded bract apices, bright red, somewhat up-curved abaxial corolla lips, and relatively long corolla beaks.

Selected References

None.

Lower Taxa

None.

0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.25 cm2.5 mm <br />0.0025 m <br />) +
Acute (?) +  and Acuminate (?) +
J. Mark Egger +, Peter F. Zika +, Barbara L. Wilson +, Richard E. Brainerd +  and Nick Otting +
(Jepson) J. M. Egger +
Castilleja spiralis +
1.5 cm15 mm <br />0.015 m <br /> (2 cm20 mm <br />0.02 m <br />) +
0.8 cm8 mm <br />0.008 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
not +, leathery +  and fleshy +
2.7 cm27 mm <br />0.027 m <br /> (2.8 cm28 mm <br />0.028 m <br />) +
yellowish green +  and green +
2-lobed +, lobed +  and tubular +
1.5 cm15 mm <br />0.015 m <br /> (2.7 cm27 mm <br />0.027 m <br />) +
taprooted;fibrous-rooted or rhizomatous +
fleshy +  and woody +
purple +, pink +, red +, orange +, yellow +, white +, green +  and pale greenish +
colored +, green +, red-violet +, red +, deep +, white +  and yellow +
0.1 cm1 mm <br />0.001 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
2 cm20 mm <br />0.02 m <br /> (3.5 cm35 mm <br />0.035 m <br />) +
oblong;narrowly spatulate +
0–3000 m. +
spreading-hairy +  and glabrous +
2 +  and 1 +
indehiscent +, septicidal +, loculicidal +  and dehiscence +
Stream banks, damp flats and seeps, over serpentine. +
stipitate-glandular +  and shaggy +
short to long +
axillary +  and terminal +
spiral +, whorled +, alternate +  and opposite +
purple +  and more or less gray +
deciduous +
basal +  and cauline +
lanceolate;linear +
2 cm20 mm <br />0.02 m <br /> (10 cm100 mm <br />0.1 m <br />) +
not fleshy +
exserted +  and abaxial +
3 +, 2 +  and 4 +
linear +  and narrowly lanceolate +
subequal +
wavy +  and plane +
tenuinucellate +  and unitegmic +
campylotropous-like +
Flowering Jun–Aug. +
oblanceolate +  and lanceolate +
20 +  and 100 +
trapezoidal +, oblong-ovoid +, pyramidal +  and ovoid +
stipitate-glandular +
Illustrated +  and Endemic +
non-secund +  and secund +
elongate +
subequal +
branched +  and unbranched +
sprawling +  and prostrate +
aerial +  and subterranean +
not fleshy +
2-lobed +  and entire +
C. minor subsp. spiralis +  and C. stenantha subsp. spiralis +
Castilleja minor var. spiralis +
Castilleja minor +
variety +
red-purple +, red +, yellow +, white +  and green +
shallow;deep +
spreading +  and strongly incurved +
reduced +
0.02 cm0.2 mm <br />2.0e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
1.6 cm16 mm <br />0.016 m <br /> (2 cm20 mm <br />0.02 m <br />) +
1.1 cm11 mm <br />0.011 m <br /> (1.6 cm16 mm <br />0.016 m <br />) +