Castilleja pallescens var. inverta

(A. Nelson & J. F. Macbride) Edwin

Leafl. W. Bot. 9: 72. 1959.

Common names: Dwarf pale or foothills paintbrush
Endemic
Basionyms: Castilleja fasciculata var. inverta A. Nelson & J. F. Macbride Bot. Gaz. 55: 381. 1913
Synonyms: C. inverta (A. Nelson & J. F. Macbride) Pennell & Ownbey
Treatment appears in FNA Volume 17. Treatment on page 638. Mentioned on page 635, 637.

Herbs 0.4–1.2 (–1.7) dm. Leaves: margins plane, 0–3 (–7) -lobed; lobes ascending-spreading. Bracts distally pale green or cream, sometimes pale yellowish or reddish purple to deep purple, lanceolate to linear-lanceolate, rigid, veins prominent, pale and contrasting with color of surfaces. Calyces purple to green with white or pink margins and veins, 15–25 (–27) mm; abaxial and adaxial clefts 7–12 mm, lateral 0.5–3 (–6) mm; lobes linear-lanceolate or narrowly triangular. Corollas subequal to calyx or beak sometimes exserted, abaxial lip sometimes partially exserted; beak adaxially whitish, buff, or pink; abaxial lip proximally green, purple, or purplish brown, distally yellow, white, or pink; teeth white, white with yellow spot proximally, cream, or pink. 2n = 24.


Phenology: Flowering May–Aug.
Habitat: Dry slopes, ridges, and plains, often with sagebrush, talus, ledges, seasonally damp meadows.
Elevation: 1500–3200 m.

Discussion

The range of var. inverta somewhat overlaps that of the similar Castilleja oresbia, but the two are separable by the characters of the key. The inflorescences of both species are usually pale straw yellow to pale green.

In Lemhi County, Idaho, primarily on alpine ridges along the summit of the Lemhi Range, plants apparently referable to var. inverta occur in large numbers. Almost all of them have deep reddish purple inflorescences, with a few rare variants of pale pink or pale yellow. This population system occurs on the border of the ranges of var. inverta and var. pallescens, and the entire complex deserves further study.

Selected References

None.

Lower Taxa

None.

0.3 cm3 mm <br />0.003 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
pink +, white +, yellow +, purplish brown +, purple +  and green +
Obtuse (?) +, Acute (?) +  and Triangular (?) +
J. Mark Egger +, Peter F. Zika +, Barbara L. Wilson +, Richard E. Brainerd +  and Nick Otting +
(A. Nelson & J. F. Macbride) Edwin +
Castilleja fasciculata var. inverta +
Pink (?) +, Buff (?) +  and Whitish (?) +
0.35 cm3.5 mm <br />0.0035 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
not +, leathery +  and fleshy +
reddish purple to deep purple;pale yellowish;cream;pale green +
lanceolate +  and linear-lanceolate +
2.5 cm25 mm <br />0.025 m <br /> (2.7 cm27 mm <br />0.027 m <br />) +
purple +  and green +
2-lobed +, lobed +  and tubular +
1.5 cm15 mm <br />0.015 m <br /> (2.5 cm25 mm <br />0.025 m <br />) +
taprooted;fibrous-rooted or rhizomatous +
fleshy +  and woody +
Dwarf pale or foothills paintbrush +
2.3 cm23 mm <br />0.023 m <br /> (2.7 cm27 mm <br />0.027 m <br />) +
purple +, pink +, red +, orange +, yellow +, white +, green +  and pale greenish +
subequal +
1.3 cm13 mm <br />0.013 m <br /> (2.3 cm23 mm <br />0.023 m <br />) +
Idaho +, Nev. +  and Oreg. +
1500–3200 m. +
spreading-hairy +  and glabrous +
2 +  and 1 +
indehiscent +, septicidal +, loculicidal +  and dehiscence +
Dry slopes, ridges, and plains, often with sagebrush, talus, ledges, seasonally damp meadows. +
stipitate-glandular +
8 cm80 mm <br />0.08 m <br /> (12 cm120 mm <br />0.12 m <br />) +
axillary +  and terminal +
spiral +, whorled +, alternate +  and opposite +
4 cm40 mm <br />0.04 m <br /> (5 cm50 mm <br />0.05 m <br />) +
green +, purple +, deep +  and purple-tinged +
deciduous +
basal +  and cauline +
linear;narrowly lanceolate +
1 cm10 mm <br />0.01 m <br /> (4 cm40 mm <br />0.04 m <br />) +
not fleshy +
ascending-spreading +
3 +, 2 +  and 4 +
triangular +  and linear-lanceolate +
subequal +
pink +  and white +
0-3(-7)-lobed +  and plane +
tenuinucellate +  and unitegmic +
campylotropous-like +
Flowering May–Aug. +
longer than deep +
expanded +
0.25 cm2.5 mm <br />0.0025 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
Leafl. W. Bot. +
20 +  and 100 +
trapezoidal +, oblong-ovoid +, pyramidal +  and ovoid +
non-secund +  and secund +
elongate +
subequal +
branched +  and unbranched +
aerial +  and subterranean +
erect +  and ascending +
not fleshy +
2-lobed +  and entire +
C. inverta +
Castilleja pallescens var. inverta +
Castilleja pallescens +
variety +
pink +, cream +, yellow spot +  and white +
shallow;deep +
erect +  and spreading +
reduced +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.35 cm3.5 mm <br />0.0035 m <br />) +
subequal +
1 cm10 mm <br />0.01 m <br /> (2 cm20 mm <br />0.02 m <br />) +