Castilleja pilosa var. steenensis

(Pennell) N. H. Holmgren in A. Cronquist et al.

Intermount. Fl. 4: 480. 1984.

Common names: Steen’s Mountain paintbrush
IllustratedEndemic
Basionyms: Castilleja steenensis Pennell Notul. Nat. Acad. Nat. Sci. Philadelphia 74: 4. 1941
Treatment appears in FNA Volume 17. Treatment on page 645. Mentioned on page 643, 646.

Stems: hairs retrorse or spreading, ± curved, stiff, 0.5–0.8 (–1) mm. Leaf lobes widely spreading. Inflorescences 2.5–10 cm; bracts proximally greenish, light dusky pink, or salmon, with margins or distal portions the same color or yellow or white, 3–5-lobed; lobes spreading to ascending, short to long, arising below mid length. Calyces (12–) 14–20 (–28) mm; abaxial and adaxial clefts 5–12 mm, 45–50% of calyx length, lateral 5–12 mm, 45–50% of calyx length; lobes linear to lanceolate. Corolla beaks 4.5–6.5 mm; abaxial lip purple or pink, sometimes pale-yellow, distally sometimes white, 3.5–6 mm, 75–100% as long as beak; teeth white or pale-yellow.


Phenology: Flowering May–Aug.
Habitat: Rocky slopes, ledges, dry meadows, sagebrush steppes, subalpine to alpine, volcanic soils.
Elevation: (1900–)2000–2900 m.

Discussion

Variety steenensis is endemic to the upper elevations of Steens Mountain in Harney County. It appears distinct when seen in its dwarfed form on the highest slopes; some transitional plants exist at lower elevations where it encounters var. pilosa. Genetic studies of this complex are in progress.

Selected References

None.

Lower Taxa

None.

"wavy-margined" is not a number.

96 +, 48 +  and 24 +
white +, pale-yellow +, pink +  and purple +
0.35 cm3.5 mm <br />0.0035 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
Rounded (?) +, Acute (?) +, Obtuse (?) +  and Acuminate (?) +
J. Mark Egger +, Peter F. Zika +, Barbara L. Wilson +, Richard E. Brainerd +  and Nick Otting +
(Pennell) N. H. Holmgren in A. Cronquist et al. +
Castilleja steenensis +
0.3 cm3 mm <br />0.003 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
not +, leathery +  and fleshy +
wavy-margined +
white;yellow;salmon;pink;salmon;pink;light dusky;greenish +
3-5-lobed +
2 cm20 mm <br />0.02 m <br /> (2.8 cm28 mm <br />0.028 m <br />) +
0.5 cm5 mm <br />0.005 m <br /> (1.2 cm12 mm <br />0.012 m <br />) +
proximally whitish +  and pale green distally green whitish pink or yellowish +
2-lobed +, lobed +  and tubular +
1.4 cm14 mm <br />0.014 m <br /> (2 cm20 mm <br />0.02 m <br />) +
taprooted;fibrous-rooted or rhizomatous +
fleshy +  and woody +
subequal +
0.2 cm2 mm <br />0.002 m <br /> (1.2 cm12 mm <br />0.012 m <br />) +
Steen’s Mountain paintbrush +
0.45 cm4.5 mm <br />0.0045 m <br /> (0.65 cm6.5 mm <br />0.0065 m <br />) +
purple +, pink +, red +, orange +, yellow +, white +, green +  and pale greenish +
1.4 cm14 mm <br />0.014 m <br /> (2.3 cm23 mm <br />0.023 m <br />) +
(1900–)2000–2900 m. +
spreading-hairy +  and glabrous +
2 +  and 1 +
indehiscent +, septicidal +, loculicidal +  and dehiscence +
Rocky slopes, ledges, dry meadows, sagebrush steppes, subalpine to alpine, volcanic soils. +
0.08 cm0.8 mm <br />8.0e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
spreading +  and retrorse +
stipitate-glandular +
wavy +  and curly +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.08 cm0.8 mm <br />8.0e-4 m <br />) +
2 cm20 mm <br />0.02 m <br /> (3.5 cm35 mm <br />0.035 m <br />) +
axillary +  and terminal +
2.5 cm25 mm <br />0.025 m <br /> (10 cm100 mm <br />0.1 m <br />) +
spiral +, whorled +, alternate +  and opposite +
5.5 cm55 mm <br />0.055 m <br /> (8 cm80 mm <br />0.08 m <br />) +
green +  and purple +
deciduous +
basal +  and cauline +
linear;lanceolate +
1 cm10 mm <br />0.01 m <br /> (5.5 cm55 mm <br />0.055 m <br />) +
not fleshy +
arising +, spreading +  and ascending +
3 +, 2 +  and 4 +
linear +  and lanceolate +
subequal +
short to long +
0-5(-7)-lobed +, plane +  and more or less wavy +
tenuinucellate +  and unitegmic +
campylotropous-like +
Flowering May–Aug. +
0.25 cm2.5 mm <br />0.0025 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
Intermount. Fl. +
20 +  and 100 +
trapezoidal +, oblong-ovoid +, pyramidal +  and ovoid +
Illustrated +  and Endemic +
non-secund +  and secund +
elongate +
subequal +
unbranched +  and branched +
aerial +  and subterranean +
ascending +  and erect +
several;many +
not fleshy +
2-lobed +  and entire +
Euchroma +  and Oncorhynchus +
Castilleja pilosa var. steenensis +
Castilleja pilosa +
variety +
pale-yellow +  and white +
shallow;deep +
reduced +
0.1 cm1 mm <br />0.001 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +