Castilleja pruinosa

Fernald

Erythea 6: 50. 1898.

Common names: Frosted or pruinose paintbrush
IllustratedEndemic
Treatment appears in FNA Volume 17. Treatment on page 647. Mentioned on page 572, 591, 611, 643, 648.

Herbs to subshrubs, perennial, (1.2–) 2–8 dm; from a woody caudex; with a taproot. Stems several to many, erect or ascending, branched or unbranched, sometimes with inconspicuous, short, leafy axillary shoots, hairs ± dense, spreading to appressed, whitish or ashy gray, long, soft, much-branched, rarely unbranched, eglandular or obscurely glandular. Leaves green, linear-lanceolate to narrowly or broadly lanceolate, 2–8 cm, not fleshy, margins plane to sometimes wavy, flat or involute, 0-lobed, sometimes 3–5-lobed distally, apex obtuse to rounded, sometimes acute; lobes sometimes divergent, ascending-spreading, linear, apex acute. Inflorescences 3–20 × 1.5–4 cm; bracts proximally greenish, distally abruptly red, sometimes yellow, salmon, or orange-red, often with a narrow yellow medial band, lanceolate, broadly lanceolate, or oblong, 0–5-lobed; lobes spreading to ascending, linear, long, arising near mid length, apex obtuse to acute. Calyces proximally green to yellow, distally and abruptly colored as in distal bracts, often with a central yellow band, 13–28 mm; abaxial clefts 5–13 mm, adaxial 8–14 mm, clefts 33–70% of calyx length, deeper than laterals, lateral 3–9 mm, 20–25% of calyx length; lobes lanceolate to triangular, apex acute to acuminate. Corollas slightly curved, 20–51 mm; tube 12–21 mm; beak long-exserted, adaxially green, yellow, or red, 12–33 mm; abaxial lip deep green, reduced, 1–2.5 mm, ca. 10% as long as beak; teeth incurved to erect, white, yellow, or green, 0.5–2 mm. 2n = 48.


Phenology: Flowering Mar–Aug.
Habitat: Dry flats, rocky slopes, talus, thickets, open forests and forest edges, usually over serpentine.
Elevation: 50–2600 m.

Discussion

Castilleja pruinosa is a common and variable paintbrush at moderate elevations in the southern half of the Cascade Range, occurring westward in Oregon and in adjacent northwestern California. It is also found in the northern Sierra Nevada. The often dense pubescence of branched hairs distinguishes it from other similar species within its range. Castilleja gleasoni was sometimes included within C. pruinosa, but C. gleasoni is likely of hybrid origin and occurs only in the San Gabriel Mountains of southern California. There are significant morphological differences between C. gleasoni and C. pruinosa. Castilleja pruinosa forms occasional hybrids with C. applegatei var. pinetorum and with C. brevilobata. In some populations in Douglas and eastern Lane counties, Oregon, C. pruinosa forms introgressive swarms with C. hispida var. hispida.

Selected References

None.

Lower Taxa

None.

"arising" is not a number.

... more about "Castilleja pruinosa"
0.5 cm5 mm <br />0.005 m <br /> (1.3 cm13 mm <br />0.013 m <br />) +
0.1 cm1 mm <br />0.001 m <br /> (0.25 cm2.5 mm <br />0.0025 m <br />) +
0.8 cm8 mm <br />0.008 m <br /> (1.4 cm14 mm <br />0.014 m <br />) +
Acute (?) +  and Acuminate (?) +
J. Mark Egger +, Peter F. Zika +, Barbara L. Wilson +, Richard E. Brainerd +  and Nick Otting +
Fernald +
Red (?) +, Yellow (?) +  and Green (?) +
long-exserted +
1.2 cm12 mm <br />0.012 m <br /> (3.3 cm33 mm <br />0.033 m <br />) +
not +, leathery +  and fleshy +
orange-red;salmon;orange-red;salmon;yellow;red;greenish +
0-5-lobed +, oblong +  and lanceolate +
colored +, proximally green +  and yellow +
2-lobed +, lobed +  and tubular +
1.3 cm13 mm <br />0.013 m <br /> (2.8 cm28 mm <br />0.028 m <br />) +
taprooted;fibrous-rooted or rhizomatous +
fleshy +  and woody +
lateral +  and deeper +
0.3 cm3 mm <br />0.003 m <br /> (0.9 cm9 mm <br />0.009 m <br />) +
Frosted or pruinose paintbrush +
purple +, pink +, red +, orange +, yellow +, white +, green +  and pale greenish +
2 cm20 mm <br />0.02 m <br /> (5.1 cm51 mm <br />0.051 m <br />) +
Calif. +  and Oreg. +
50–2600 m. +
spreading-hairy +  and glabrous +
2 +  and 1 +
indehiscent +, septicidal +, loculicidal +  and dehiscence +
Dry flats, rocky slopes, talus, thickets, open forests and forest edges, usually over serpentine. +
glandular +, eglandular +, unbranched +  and much-branched +
ashy gray +  and whitish +
spreading +  and appressed +
stipitate-glandular +
axillary +  and terminal +
spiral +, whorled +, alternate +  and opposite +
deciduous +
basal +  and cauline +
linear-lanceolate;narrowly or broadly lanceolate +
2 cm20 mm <br />0.02 m <br /> (8 cm80 mm <br />0.08 m <br />) +
not fleshy +
spreading +  and ascending +
3 +, 2 +  and 4 +
lanceolate +  and triangular +
subequal +
3-5-lobed +, 0-lobed +, involute +, flat +, wavy +  and plane +
tenuinucellate +  and unitegmic +
campylotropous-like +
Flowering Mar–Aug. +
20 +  and 100 +
trapezoidal +, oblong-ovoid +, pyramidal +  and ovoid +
Illustrated +  and Endemic +
non-secund +  and secund +
elongate +
subequal +
unbranched +  and branched +
sprawling +  and prostrate +
aerial +  and subterranean +
ascending +  and erect +
several;many +
not fleshy +
2-lobed +  and entire +
Euchroma +  and Oncorhynchus +
Castilleja pruinosa +
Castilleja +
species +
green +, yellow +  and white +
shallow;deep +
incurved +  and erect +
reduced +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
1.2 cm12 mm <br />0.012 m <br /> (2.1 cm21 mm <br />0.021 m <br />) +