Ceratodon heterophyllus

Kindberg

Ottawa Naturalist 5: 179. 1892,.

Synonyms: Ceratodon purpureus var. obtusifolius Limpricht Ceratodon purpureus var. rotundifolius Berggren
Treatment appears in FNA Volume 27. Treatment on page 446. Mentioned on page 445.

Plants in dense tufts or mats, green to dark green to brownish green. Stems 0.3–1 cm. Leaves loosely imbricate, somewhat contorted when dry, ovate to ovatelanceolate, concave and often somewhat cucullate, (0.35–) 0.5–0.9 (–1.5) mm, margins plane to weakly recurved, often only at midleaf, and entire to, rarely, weakly toothed distally, apices obtuse; most costae ending before apex; medial laminal cells (9–) 12–16 (–22) µm, thin-walled to somewhat thickened. Seta (0.9–) 1.5 (–2.5) cm, red. Capsule ovate-cylindric to, occasionally, ovate, (0.9–) 1–1.2 (–1.5) mm, strongly sulcate when dry; peristome segments free at their nodes, finely papillose, dark red with pale borders to, rarely, completely pale. Spores (18–) 19–21 (–22) µm.


Phenology: Capsules mature early summer–late fall.
Habitat: Sandy, usually moist soil, Arctic habitats
Elevation: low elevations (0-200 m)

Distribution

V27 641-distribution-map.gif

Greenland, N.W.T., Yukon, Alaska, Eurasia

Discussion

In Northwest Territories, Ceratodon heterophyllus is found only on Prince Patrick Island.

Ceratodon heterophyllus has also been treated as C. purpureus forma heterophyllus (Kindberg) Britton, and R. R. Ireland (1980) considered it to be a variety of C. purpureus, but J. S. Burley and N. M. Pritchard (1990) provided ample evidence that it is a distinct species. The broadly ovate leaves of C. heterophyllus grade into some alpine variations of C. purpureus.

Lower Taxa

None.
... more about "Ceratodon heterophyllus"
forming a marginal border +  and not differentiated +
Terry T. McIntosh +
Kindberg +
finely papillose;spinulose-papillose +
cucullate +
0.12 cm1.2 mm <br />0.0012 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
dark red;reddish or purplish brown +
cleistocarpous +
erect;inclined or horizontal +
ovate +  and ovate-cylindric +
0.1 cm1 mm <br />0.001 m <br /> (0.12 cm1.2 mm <br />0.0012 m <br />) +
deciduous +
subpercurrent +
shortrectangular +  and isodiametric +
Greenland +, N.W.T. +, Yukon +, Alaska +  and Eurasia +
low elevations (0-200 m) +
Sandy, usually moist soil, Arctic habitats +
0.09 cm0.9 mm <br />9.0e-4 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
straight +
cucullate +, concave +, ovate +  and ovatelanceolate +
quadrate +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.09 cm0.9 mm <br />9.0e-4 m <br />) +
smooth +  and serrate +
toothed;plane +
dioicous +  and sexual +
rounded +  and angled +
shortrectangular +
conic +  and long-conic +
abruptly subulate +  and gradually acuminate +
with a longer , broader sheathing base +  and differentiated +
long-acuminate +, short-acuminate +, concave +  and ovate +
on short branches +  and axillary +
pale +  and dark red +
Capsules mature early summer–late fall. +
well-developed +
filamentous +
Ottawa Naturalist +
ireland1980a +
papillose +
free;united +
1.5 cm15 mm <br />0.015 m <br /> (2.5 cm25 mm <br />0.025 m <br />) +
elongate +
1.5 cm15 mm <br />0.015 m <br /> (?) +
globose +
0.3 cm3 mm <br />0.003 m <br /> (1 cm10 mm <br />0.01 m <br />) +
Ceratodon purpureus var. obtusifolius +  and Ceratodon purpureus var. rotundifolius +
Ceratodon heterophyllus +
Ceratodon +
species +
specialized +
synoicous +, paroicous +, autoicous +, dioicous +  and sexual +
green +  and dark green +
female +  and male +
small +  and medium-sized +