Chaenactis glabriuscula var. heterocarpha

(Torrey & A. Gray) H. M. Hall

Univ. Calif. Publ. Bot. 3: 190. 1907.

Common names: Inner Coast Range chaenactis
Endemic
Basionym: Chaenactis heterocarpha Torrey & A. Gray Mem. Amer. Acad. Arts, n. s. 4: 98. 1849
Synonyms: Chaenactis glabriuscula var. gracilenta (Greene) D. D. Keck Chaenactis heterocarpha var. tanacetifolia (A. Gray) A. Gray Chaenactis tanacetifolia Chaenactis tanacetifolia var. gracilenta (Greene) Stockwell
Treatment appears in FNA Volume 21. Treatment on page 412. Mentioned on page 409, 411.

Plants 6–40 cm; proximal indument grayish, arachnoid to sparsely lanuginose. Stems mostly 1–5, erect to spreading; branches proximal and, often, distal. Leaves basal (often withering) and ± cauline, 1–6 (–10) cm; largest blades ± 3-dimensional, scarcely succulent, usually 2-pinnately lobed; primary lobes 2–7 pairs, remote to ± congested, ultimate lobes ± involute and/or twisted. Heads mostly (1–) 2–5 per stem. Peduncles 2–7 (–9) cm. Involucres ± hemispheric to broadly cylindric. Phyllaries: longest 6–10 × 1.5–2.5 mm; outer glabrescent to sericeous and, often, ± glandular in fruit. Florets: inner corollas 4.5–7 mm. Cypselae 4–9 mm; pappi of (5–) 8 scales in 2, abruptly unequal series, longest scales mostly 2–7 mm, lengths 0.4–0.9 (–1) times corollas. 2n = 12.


Phenology: Flowering Mar–Jun.
Habitat: Slopes, ridges, mostly over serpentine, openings in chaparral, woodlands
Elevation: 100–1500 m

Discussion

Variety heterocarpha is known mainly from west-central and northwestern California. It intergrades to minor degrees with vars. megacephala, lanosa, and glabriuscula at its southern limits. Most of the specimens from the Tehachapi Mountains region assigned to this taxon appear to represent hybrids between C. xantiana and vars. megacephala or glabriuscula (see sectional discussion). Chaenactis tanacetifolia is an occasional form with leaves mostly basal, persistent, and densely arachnoid, and lobes more congested; its taxonomic significance remains to be demonstrated.

Selected References

None.

Lower Taxa

None.

"-0.9(-1)timescorollas" is not declared as a valid unit of measurement for this property.

not tailed +, rounded +  and obtuse +
connate +  and distinct +
subequal +
usually ovate +  and lanceolate +
usually deltate +  and lanceolate +
paniculiform +, cymiform +  and corymbiform +
indeterminate +  and determinate +
James D. Morefield +
(Torrey & A. Gray) H. M. Hall +
decurrent +
Chaenactis heterocarpha +
compound +  and simple +
lobed;ovate;oblanceolate;ovate;oblanceolate;linear;deltate +
winged;ribbed;winged;ribbed +
tuberculate +  and rugose +
barbellulate +  and barbellate +
connate +  and distinct +
Inner Coast Range chaenactis +
actinomorphic +
stipitate-glandular +  and hairy +
0.4 cm4 mm <br />0.004 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
fertile +  and bisexual +
fertile +  and bisexual +
100–1500 m +
strigose +  and densely sericeous +
scabrellous +
pistillate +, staminate +  and neuter +
winged +  and beaked +
dispersed +
campanulate +  and cylindric +
Slopes, ridges, mostly over serpentine, openings in chaparral, woodlands +
indeterminate +
homogamous +  and heterogamous +
each +  and sessile +
0.45 cm4.5 mm <br />0.0045 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
0.5 cm5 mm <br />0.005 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
less hemispheric;broadly cylindric +
6 cm60 mm <br />0.06 m <br /> (10 cm100 mm <br />0.1 m <br />) +
cauline +  and basal +
1 cm10 mm <br />0.01 m <br /> (?) +  and 6 cm60 mm <br />0.06 m <br /> (?) +
deltate +  and more or less lanceolate +
0.1 cm1 mm <br />0.001 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
0.2 cm2 mm <br />0.002 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
dentate to pinnatifid or palmatifid +  and entire +
glabrescent +  and sericeous +
acute +  and blunt +
2-carpellate +
persistent +
coroniform +
7 cm70 mm <br />0.07 m <br /> (9 cm90 mm <br />0.09 m <br />) +
erect +  and ascending +
glabrescent +, lanuginose +, arachnoid-sericeous +, villous +  and stipitate-glandular +
2 cm20 mm <br />0.02 m <br /> (7 cm70 mm <br />0.07 m <br />) +
Flowering Mar–Jun. +
0.6 cm6 mm <br />0.006 m <br /> (1 cm10 mm <br />0.01 m <br />) +
5 +  and 21 +
0.45 cm4.5 mm <br />0.0045 m <br /> (1 cm10 mm <br />0.01 m <br />) +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.25 cm2.5 mm <br />0.0025 m <br />) +
remote +  and more or less congested +
arachnoid +  and sparsely lanuginose +
Univ. Calif. Publ. Bot. +
bearing subulate enations +, hairy +  and bristly +
knobby;convex;more or less flat +
distinct +
unequal +
exalbuminous +
unequal +
5 (?) +  and 12 (?) +
erect +  and spreading +
1 +  and 5 +
appendaged +  and truncate +
hirsutulous +  and glabrous +
papillate +
Chaenactis glabriuscula var. gracilenta +, Chaenactis heterocarpha var. tanacetifolia +, Chaenactis tanacetifolia +  and Chaenactis tanacetifolia var. gracilenta +
Chaenactis glabriuscula var. heterocarpha +
Chaenactis glabriuscula +
variety +
cylindric +
80 cm800 mm <br />0.8 m <br /> (300 cm3,000 mm <br />3 m <br />) +
1 cm10 mm <br />0.01 m <br /> (80 cm800 mm <br />0.8 m <br />) +
toothed +  and entire +
70 cm700 mm <br />0.7 m <br /> (200 cm2,000 mm <br />2 m <br />) +
5 cm50 mm <br />0.05 m <br /> (70 cm700 mm <br />0.7 m <br />) +
80 cm800 mm <br />0.8 m <br /> (300 cm3,000 mm <br />3 m <br />) +