Cheilanthes cooperae

D. C. Eaton

Bull. Torrey Bot. Club 6: 33. 1875.

Common names: Mrs. Cooper's lip fern
Endemic
Treatment appears in FNA Volume 2.

Stems compact to short-creeping, usually 4–8 mm diam.; scales brown at base and darker toward tip, linear-subulate, straight to slightly contorted, loosely appressed, persistent. Leaves clustered, 5–30 cm; vernation circinate. Petiole dark-brown, flattened or slightly grooved distally on adaxial surface. Blade linear-oblong to lanceolate-ovate, 3-pinnate at base, 1.5–5 cm wide; rachis flattened or slightly grooved adaxially, lacking scales, with monomorphic pubescence. Pinnae not articulate, dark color of stalk continuing into pinna base, basal pair slightly smaller than adjacent pair, ± equilateral, appearing hirsute adaxially. Costae green adaxially for most of length; abaxial scales absent. Ultimate segments linear-oblong to ovate, not beadlike, the largest 3–5 mm, abaxially and adaxially hirsute with long, strongly flattened hairs. False indusia marginal, weakly differentiated, 0.05–0.25 mm wide. Sori usually discontinuous, concentrated on apical and lateral lobes. Sporangia containing 64 spores. 2n = 60.


Phenology: Sporulating late spring–summer.
Habitat: Calcareous cliffs and ledges, usually on limestone
Elevation: 100–700 m

Discussion

Although scattered throughout much of California, Cheilanthes cooperae is apparently rare and quite localized. It appears to be most closely related to C. viscida, from which it differs in lacking glandular pubescence. The ranges of the two species do not overlap, and they seem amply distinct.

Selected References

None.

Lower Taxa

None.

... more about "Cheilanthes cooperae"
Michael D. Windham +  and Eric W. Rabe +
D. C. Eaton +
slightly smaller than adjacent pair +
cuneate +, truncate +  and rounded +
scaly rarely glabrous adaxially pubescent +  and glabrous +
linear-oblong +  and lanceolate-ovate +
1.5cm;5cm +
Mrs. Cooper's lip fern +
100–700 m +
0.005 cm0.05 mm <br />5.0e-5 m <br /> (0.025 cm0.25 mm <br />2.5e-4 m <br />) +
yellow +  and white +
obcordate;reniform +
Calcareous cliffs and ledges, usually on limestone +
flattened +
monomorphic +
clustered;widely scattered +
recurved +  and reflexed +
noncircinate +  and circinate +
5 cm50 mm <br />0.05 m <br /> (30 cm300 mm <br />0.3 m <br />) +
circinate +
denticulate +  and entire +
glabrous +, scaly +  and pubescent +
flattened +
Sporulating late spring–summer. +
not articulate +
Bull. Torrey Bot. Club +
flattened +
darker;brown +
straight +
persistent +
linear-subulate +
yellowish +, brown +  and black or gray +
cristate +  and tetrahedral-globose +
dark colored +
0.4 cm4 mm <br />0.004 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
ascending +  and horizontal +
Cheilanthes cooperae +
Cheilanthes +
species +
0.3 cm3 mm <br />0.003 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
0 cm0 mm <br />0 m <br /> (?) +  and 0.4 cm4 mm <br />0.004 m <br /> (?) +
crescent--shaped +  and roundish +
branched +  and anastomosing +