Cheilanthes intertexta

(Maxon) Maxon in Abrams

in Abrams,Ill. Fl. Pacific States 1: 28. 1923.

Common names: Coastal lip fern
Basionym: Cheilanthes covillei subsp. intertexta Maxon Proc. Biol. Soc. Wash. 31: 149. 1918
Treatment appears in FNA Volume 2.

Stems short-creeping, usually 3–7 mm diam.; scales usually bicolored, with broad, well-defined, dark, central stripe and narrow, light-brown margins, linear-lanceolate, straight to slightly contorted, strongly appressed, persistent. Leaves clustered, 4–25 cm; vernation noncircinate. Petiole dark-brown, rounded adaxially. Blade lanceolate to ovate-deltate, usually 3-pinnate at base, 1–4 cm wide; rachis rounded adaxially, with scattered scales and sparse monomorphic pubescence. Pinnae not articulate, dark color of stalk continuing into pinna base, basal pair not conspicuously larger than adjacent pair, usually equilateral, appearing glabrous to sparsely pubescent adaxially. Costae green adaxially for most of length; abaxial scales multiseriate, ovatelanceolate, deeply cordate at base, with overlapping basal lobes, conspicuous, the longest 0.4–1 mm wide, imbricate, often concealing ultimate segments, long-ciliate, cilia usually confined to proximal 1/2. Ultimate segments oblong to ovate, beadlike, the largest 1–3 mm, abaxially densely covered with branched hairs and small, ciliate scales, adaxially with scattered branched hairs or glabrescent. False indusia marginal, weakly differentiated, 0.05–0.25 mm wide. Sori ± continuous around segment margins. Sporangia containing 64 spores.

Phenology: Sporulating late spring–fall.
Habitat: Rocky slopes and ledges, usually on igneous substrates
Elevation: 500–2800 m


Preliminary isozyme analyses support D. B. Lellinger's (1985) suggestion that Cheilanthes intertexta is a fertile allotetraploid hybrid between C. gracillima and C. covillei. It is morphologically most similar to the latter parent (see comments under C. covillei), but it is occasionally confused with C. gracillima, with which it apparently hybridizes to form sterile intermediates that have been called C. gracillima var. aberrans M. E. Jones. Cheilanthes intertexta may also be confused with C. clevelandii, with which it is partially sympatric. In addition to the characters given in the key, C. intertexta is distinguished from closely related sexual species by having larger spores averaging more than 55 µm in diameter.

Selected References


Lower Taxa

... more about "Cheilanthes intertexta"
cordate +  and ovatelanceolate +
0.04 cm0.4 mm <br />4.0e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
Michael D. Windham +  and Eric W. Rabe +
(Maxon) Maxon in Abrams +
glabrous +  and sparsely pubescent +
not conspicuously larger than adjacent pair +
cuneate +, truncate +  and rounded +
Cheilanthes covillei subsp. intertexta +
scaly rarely glabrous adaxially pubescent +  and glabrous +
lanceolate +  and ovate-deltate +
1cm;4cm +
Coastal lip fern +
Calif. +, Nev. +  and Oreg. +
500–2800 m +
0.005 cm0.05 mm <br />5.0e-5 m <br /> (0.025 cm0.25 mm <br />2.5e-4 m <br />) +
yellow +  and white +
obcordate;reniform +
Rocky slopes and ledges, usually on igneous substrates +
monomorphic +
clustered;widely scattered +
recurved +  and reflexed +
noncircinate +  and circinate +
4 cm40 mm <br />0.04 m <br /> (25 cm250 mm <br />0.25 m <br />) +
light-brown +
persistent +
linear-lanceolate +
glabrous +, scaly +  and pubescent +
Sporulating late spring–fall. +
not articulate +
in Abrams,Ill. Fl. Pacific States +
monomorphic +
persistent +
linear-subulate +  and ovatelanceolate +
yellowish +, brown +  and black or gray +
cristate +  and tetrahedral-globose +
dark colored +
0.3 cm3 mm <br />0.003 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
ascending +  and horizontal +
Cheilanthes intertexta +
Cheilanthes +
species +
0.1 cm1 mm <br />0.001 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
0 cm0 mm <br />0 m <br /> (?) +  and 0.4 cm4 mm <br />0.004 m <br /> (?) +
crescent--shaped +  and roundish +
branched +  and anastomosing +