Cheilanthes newberryi

(D. C. Eaton) Domin

Biblioth. Bot. 20(85): 133. 1915.

Common names: Newberry's lip fern
Basionyms: Notholaena newberryi D. C. Eaton Bull. Torrey Bot. Club 4: 12. 1873
Treatment appears in FNA Volume 2.

Stems short-creeping to compact, usually 3–5 mm diam.; scales uniformly dark-brown to black or with light-brown, ephemeral tip, linear-subulate, slightly contorted, strongly appressed, persistent. Leaves clustered, 5–30 cm; vernation noncircinate. Petiole dark-brown, rounded adaxially. Blade lanceolate to linear-oblong, 2-pinnate-pinnatifid to 3-pinnate at base, 1.5–5 cm wide; rachis rounded adaxially, lacking scales, with dense monomorphic pubescence. Pinnae not articulate, dark color of stalk continuing into pinna base, basal pair slightly smaller than adjacent pair, ± equilateral, proximal basiscopic pinnules slightly enlarged, appearing tomentose adaxially. Costae green or straw-colored adaxially for most of length; abaxial scales absent. Ultimate segments oblong to lanceolate, not beadlike, the largest 3–5 mm, abaxially densely woolly with very fine, cobwebby hairs, tomentose adaxially. False indusia absent. Sori usually discontinuous, concentrated on small apical and lateral lobes. Sporangia containing 64 or 32 spores. 2n = 60.


Phenology: Sporulating late spring–summer.
Habitat: Rocky cliffs and slopes, usually on igneous substrates
Elevation: 0–1200 m

Distribution

V2 36-distribution-map.gif

Calif., Mexico in Baja California

Discussion

As with Cheilanthes bonariensis, C. newberryi has traditionally been assigned to Notholaena; R. M. Tryon and A. F. Tryon (1982) argued for its placement in Cheilanthes. Hybridization of C. newberryi with C. covillei to form C. × fibrillosa supports placement in Cheilanthes. R. M. Tryon (1956) reported variations in spore number per sporangium in C. newberryi that may indicate cytologic variability within the species.

Selected References

None.

Lower Taxa

None.

"fine" is not a number.

... more about "Cheilanthes newberryi"
Michael D. Windham +  and Eric W. Rabe +
(D. C. Eaton) Domin +
slightly smaller +
cuneate +, truncate +  and rounded +
Notholaena newberryi +
scaly rarely glabrous adaxially pubescent +  and glabrous +
lanceolate +  and linear-oblong 2-pinnate-pinnatifid +
1.5cm;5cm +
Newberry's lip fern +
straw-colored +  and green +
Calif. +  and Mexico in Baja California +
0–1200 m +
rarely +  and marginal +
yellow +  and white +
obcordate;reniform +
Rocky cliffs and slopes, usually on igneous substrates +
monomorphic +
clustered;widely scattered +
recurved +  and reflexed +
noncircinate +  and circinate +
5 cm50 mm <br />0.05 m <br /> (30 cm300 mm <br />0.3 m <br />) +
denticulate +  and entire +
glabrous +, scaly +  and pubescent +
Sporulating late spring–summer. +
not articulate +
enlarged +
Biblioth. Bot. +
with light-brown , ephemeral tip;uniformly dark-brown;black +
persistent +
linear-subulate +
32 (?) +  and 64 (?) +
yellowish +, brown +  and black or gray +
1 +, 32 +  and 64 +
cristate +  and tetrahedral-globose +
dark colored +
0.3 cm3 mm <br />0.003 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
ascending +  and horizontal +
Cheilanthes newberryi +
Cheilanthes +
species +
light-brown +
ephemeral +
0.3 cm3 mm <br />0.003 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
0 cm0 mm <br />0 m <br /> (?) +  and 0.4 cm4 mm <br />0.004 m <br /> (?) +
crescent--shaped +  and roundish +
branched +  and anastomosing +