Cirsium douglasii var. breweri

(A. Gray) D. J. Keil & C. E. Turner

Phytologia 73: 313. 1992.

Common names: Brewer’s thistle
IllustratedEndemic
Basionym: Cnicus breweri A. Gray Proc. Amer. Acad. Arts 10: 43. 1874
Synonyms: Cirsium breweri (A. Gray) Jepson Cirsium douglasii var. canescens (Petrak) J. T. Howell
Treatment appears in FNA Volume 19. Treatment on page 133.

Plants to 250 cm. Cauline leaves usually unlobed or shallowly lobed, spines usually less than 2–7 mm. Heads mostly 2–3 cm. 2n = 30.


Phenology: Flowering summer (Jun–Aug).
Habitat: Wet soil, streams, fens, marshes, springs in montane coniferous forest areas, often on serpentine
Elevation: 1200–1900 m

Discussion

Variety breweri grows from the eastern North Coast Ranges of California and the Klamath and Siskiyou ranges east to the northen Sierra Nevada.

Selected References

None.

Lower Taxa

None.

"fine" is not a number."[" is not declared as a valid unit of measurement for this property."]" is not declared as a valid unit of measurement for this property."-7mm" is not declared as a valid unit of measurement for this property.

short-tailed +
spineless +  and twisted +
dentate +, entire +  and spine-tipped +
erect +  and spreading +
innermost +
scarious +
not differentiated +
paniculiform +
indeterminate +  and determinate +
David J. Keil +
(A. Gray) D. J. Keil & C. E. Turner +
auriculate-clasping +
decurrent +
Cnicus breweri +
compound +  and simple +
10 cm100 mm <br />0.1 m <br /> (60 cm600 mm <br />0.6 m <br />) +
unlobed or;shallowly deeply pinnatifid +
2cm;15cm +
10-nerved or 20-nerved +  and rugose +
ascending;spreading +
tawny +  and white +
? (?) +  and ? (?) +
lobed +  and unlobed +
Brewer’s thistle +
actinomorphic +
1.8 cm18 mm <br />0.018 m <br /> (2.1 cm21 mm <br />0.021 m <br />) +
dark-brown +  and black +
compressed +  and ovoid +
fertile +  and bisexual +
Calif. +, Nev. +  and Oreg. +
1200–1900 m +
glabrate +  and gray-tomentose +
pistillate +, staminate +  and neuter +
winged +  and beaked +
dispersed +
Wet soil, streams, fens, marshes, springs in montane coniferous forest areas, often on serpentine +
indeterminate +
2 cm20 mm <br />0.02 m <br /> (3 cm30 mm <br />0.03 m <br />) +
each +  and sessile +
2 cm20 mm <br />0.02 m <br /> (4.5 cm45 mm <br />0.045 m <br />) +
glabrate +  and arachnoid +
ovoid;hemispheric +
1.5 cm15 mm <br />0.015 m <br /> (3 cm30 mm <br />0.03 m <br />) +
sessile +  and petiolate +
cauline +  and basal +
bristle-tipped +
ascending;spreading +
lobed +, dentate +, lanceolate +  and ovate-triangular +
0.2 cm2 mm <br />0.002 m <br /> (3 cm30 mm <br />0.03 m <br />) +
entire +  and dentate +
usually lobed +  and dissected +
2-carpellate +
persistent +
1.5 cm15 mm <br />0.015 m <br /> (2 cm20 mm <br />0.02 m <br />) +
dark purple +
4 cm40 mm <br />0.04 m <br /> (8 cm80 mm <br />0.08 m <br />) +
0 cm0 mm <br />0 m <br /> (4 cm40 mm <br />0.04 m <br />) +
Flowering summer (Jun–Aug). +
middle +  and outer +
weakly to strongly +
papillate +  and smooth +
bearing subulate enations +, hairy +  and bristly +
flat;convex +
linear to elliptic +
glutinous +
tawny;white +
setiform +  and plumose +
exalbuminous +
subequal +
Illustrated +  and Endemic +
spiny-winged +, simple +  and branched +
ascending +  and erect +
1 +  and several +
appendaged +  and truncate +
dilated +  and swollen +
enlarged +
0.3 cm3 mm <br />0.003 m <br /> (0.45 cm4.5 mm <br />0.0045 m <br />) +
Cirsium breweri +  and Cirsium douglasii var. canescens +
Cirsium douglasii var. breweri +
Cirsium douglasii +
variety +
cylindric +
expanded +
0.5 cm5 mm <br />0.005 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
bristly-dentate to coarsely +
slender +
0.8 cm8 mm <br />0.008 m <br /> (0.9 cm9 mm <br />0.009 m <br />) +
perennial +, short-lived +  and biennial +
60 cm600 mm <br />0.6 m <br /> (250 cm2,500 mm <br />2.5 m <br />) +
1 cm10 mm <br />0.01 m <br /> (3 cm30 mm <br />0.03 m <br />) +