Clarkia purpurea subsp. quadrivulnera

(Douglas ex Lindley) H. Lewis & M. E. Lewis

Univ. Calif. Publ. Bot. 20: 305. 1955.

Common names: Purple godetia
Basionym: Oenothera quadrivulnera Douglas ex Lindley Bot. Reg. 13: plate 1119. 1828
Synonyms: Clarkia quadrivulnera (Douglas ex Lindley) A. Nelson & J. F. Macbride Godetia purpurea var. parviflora (S. Watson) C. L. Hitchcock G. quadrivulnera (Douglas ex Lindley) Spach G. quadrivulnera var. vacensis Jepson
Treatment appears in FNA Volume 10.

Leaf-blades linear to lanceolate, 1.5–5 cm. Inflorescences usually open racemes. Flowers: petals lavender to purple or dark wine-red, often with purple spot near middle or distally, 9–14 mm; stigma not exserted beyond anthers. 2n = 52.


Phenology: Flowering Apr–Aug.
Habitat: Open grassy or shrubby places.
Elevation: 0–500 m.

Distribution

B.C., Ariz., Calif., Oreg., Wash., Mexico (Baja California)

Discussion

Subspecies quadrivulnera is extremely widespread from Baja California in Mexico north through California, Oregon, and Washington to the islands of extreme southwestern British Columbia.

Subspecies quadrivulnera is diverse in petal color and color pattern but all of the populations have relatively small flowers that are primarily self-pollinated, usually with pollen in contact with the stigma at the time the flowers open.

Selected References

None.

Lower Taxa

None.
Harlan Lewis† +  and Peter C. Hoch +
(Douglas ex Lindley) H. Lewis & M. E. Lewis +
straight +
prostrate +  and decumbent +
0.9 cm9 mm <br />0.009 m <br /> (2.5 cm25 mm <br />0.025 m <br />) +
Oenothera quadrivulnera +
0 cm0 mm <br />0 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
indehiscent +  and loculicidal +
1.5 cm15 mm <br />0.015 m <br /> (5 cm50 mm <br />0.05 m <br />) +
linear +  and lanceolate +
denticulate +  and entire +
narrowly lanceolate +  and elliptic or ovate +
1.5 cm15 mm <br />0.015 m <br /> (7 cm70 mm <br />0.07 m <br />) +
curved +  and straight +
indehiscent +  and loculicidal +
terete +, 4-angled +, clavate +, fusiform +, cylindrical +  and elongate +
1 cm10 mm <br />0.01 m <br /> (3 cm30 mm <br />0.03 m <br />) +
elongated +
inconspicuous +
Purple godetia +
bowl--shaped +
0.2 cm2 mm <br />0.002 m <br /> (?) +
B.C. +, Ariz. +, Calif. +, Oreg. +, Wash. +  and Mexico (Baja California) +
0–500 m. +
basifixed +  and versatile +
0.2 cm2 mm <br />0.002 m <br /> (1 cm10 mm <br />0.01 m <br />) +
actinomorphic +
protandrous +  and bisexual +
pedicellate +  and sessile +
curved +  and straight +
indehiscent +  and loculicidal +
terete +, 4-angled +, clavate +, fusiform +, cylindrical +  and elongate +
Open grassy or shrubby places. +
unicellular-papillose +
deciduous +
purple spot +
numerous +  and 1 +
lavender +  and purple or dark wine-red +
unlobed +, obovate +  and obdeltate +
0.9 cm9 mm <br />0.009 m <br /> (1.4 cm14 mm <br />0.014 m <br />) +
0 cm0 mm <br />0 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
Flowering Apr–Aug. +
red +, lavender +, blue-gray +, yellow +  and cream +
Univ. Calif. Publ. Bot. +
basal +  and cauline +
several +  and 1 +
clavate +  and globose +
gray +  and brown +
few +  and numerous +
sculptured +  and smooth +
0.1 cm1 mm <br />0.001 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
pink +  and purplish red +
deciduous +
in pairs +  and reflexed +
parenchymatous +
parietal +, axile +  and placentation +
1 +, 4 +  and 2 +
unequal +
aquatic +, amphibious +  and terrestrial +
subequal +
2 times as many or as many as sepals +
unbranched +  and sparsely branched +
decumbent +  and prostrate +
rarely +  and erect +
glaucous or +  and sparsely densely puberulent +
slender +  and stout +
0 cm0 mm <br />0 m <br /> (100 cm1,000 mm <br />1 m <br />) +
petiolate +  and sessile +
intrapetiolar +
aquatic +, amphibious +  and terrestrial +
Clarkia quadrivulnera +, Godetia purpurea var. parviflora +, G. quadrivulnera +  and G. quadrivulnera var. vacensis +
Clarkia purpurea subsp. quadrivulnera +
Clarkia purpurea +
subspecies +
3(-5)-aperturate +
short-acute +
0.9 cm9 mm <br />0.009 m <br /> (2.5 cm25 mm <br />0.025 m <br />) +
smooth +  and erose +
papillate +