Comandra umbellata subsp. pallida

(A. de Candolle) Piehl

Mem. Torrey Bot. Club 22(1): 70. 1965.

Basionym: Comandra pallida A. de Candolle in A. P. de Candolle and A. L. P. P. de Candolle Prodr. 14: 636. 1857
Synonyms: C. umbellata var. pallida (A. de Candolle) M. E. Jones
Treatment appears in FNA Volume 12. Treatment on page 410. Mentioned on page 409, 411.

Herbs, 5–33 cm. Rhizomes: cortex blue, drying blackish. Aerial stems often much branched; proximal portions not overwintering. Leaves: blade green to grayish green, glaucous, linear to lanceolate, elliptic, or ovate, 0.9–4.2 cm, thick, firm, becoming ± succulent, base attenuate to cuneate, margins not revolute, apex attenuate to acute, often mucronulate or apiculate; midrib obscure or sometimes conspicuous or protruding on abaxial surface, lateral-veins obscure. Pedicels 0–1.6 mm. Flowers funnel-shaped to rotate; petals oblong to lanceolate-oblong, 2.5–5 mm; anthers 0.7 mm. Pseudodrupes purplish brown to red, glaucous, subglobose to ovate, 5.5–9 mm, slightly roughened. 2n = 52.


Phenology: Flowering Mar–Jul.
Habitat: Open areas, sandy and rocky slopes, sagebrush communities, coniferous and deciduous forests, deserts.
Elevation: 150–2800 m.

Distribution

V12 749-distribution-map.jpg

Alta., B.C., Man., N.W.T., Sask., Yukon, Ariz., Colo., Idaho, Kans., Mont., Nebr., Nev., N.Mex., N.Dak., Okla., Oreg., S.Dak., Tex., Utah, Wash., Wyo., Mexico (Chihuahua), Mexico (Coahuila)

Discussion

As with the other subspecies, subsp. pallida is morphologically variable, this likely related to its diverse ecological habitats. Clinal variation and intergradation with the other subspecies that flank its eastern and western boundaries may also play a role (M. A. Piehl 1965). Intermediates between subsp. pallida and subsp. californica occur in Arizona, New Mexico, Nevada, Oregon, and Washington. Ecotypic variation apparently has a genetic component, as demonstrated by common garden experiments with seeds and vegetative propagules (Piehl). One extreme but widespread form is highly branched basally and has dimorphic leaves, with the proximal blades linear and the distal blades lanceolate or elliptic.

Selected References

None.

Lower Taxa

None.
0.7 cm7 mm <br />0.007 m <br /> (?) +
Apiculate (?) +, Mucronulate (?) +, Attenuate (?) +  and Acute (?) +
Daniel L. Nickrent +
(A. de Candolle) Piehl +
attenuate;cuneate +
Comandra pallida +
light green;grayish or bluish green +
0.7 cm7 mm <br />0.007 m <br /> (5.3 cm53 mm <br />0.053 m <br />) +
ovate +, elliptic +  and lanceolate +
green;grayish green +
linear +  and lanceolate elliptic or ovate +
0.9 cm9 mm <br />0.009 m <br /> (4.2 cm42 mm <br />0.042 m <br />) +
? (?) +  and ? (?) +
blackish +  and blue +
exfoliating +
papery +  and corky +
Alta. +, B.C. +, Man. +, N.W.T. +, Sask. +, Yukon +, Ariz. +, Colo. +, Idaho +, Kans. +, Mont. +, Nebr. +, Nev. +, N.Mex. +, N.Dak. +, Okla. +, Oreg. +, S.Dak. +, Tex. +, Utah +, Wash. +, Wyo. +, Mexico (Chihuahua) +  and Mexico (Coahuila) +
150–2800 m. +
persistent +
Open areas, sandy and rocky slopes, sagebrush communities, coniferous and deciduous forests, deserts. +
free +  and adnate +
corymblike +  and paniclelike +
protruding +, conspicuous +, sometimes +  and obscure +
0 cm0 mm <br />0 m <br /> (0.16 cm1.6 mm <br />0.0016 m <br />) +
persistent +
free +, distinct +  and connate +
0.25 cm2.5 mm <br />0.0025 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
Flowering Mar–Jul. +
not overwintering +
purplish brown +  and red +
0.55 cm5.5 mm <br />0.0055 m <br /> (0.9 cm9 mm <br />0.009 m <br />) +
Mem. Torrey Bot. Club +
white;beige or blue +
deciduous +
andromonoecious +  and synoecious +
Illustrated +
filiform +
C. umbellata var. pallida +
Comandra umbellata subsp. pallida +
Comandra umbellata +
subspecies +