Dicranum fuscescens var. flexicaule

(Bridel) Wilson

Bryol. Brit., 77. 1855,.

Basionym: Dicranum flexicaule Bryol. Univ. 1: 421. 1826
Treatment appears in FNA Volume 27. Treatment on page 413. Mentioned on page 412.

Plants in loose tufts. Stems 8-18 cm, scarcely tomentose. Leaves strongly falcate-secund, somewhat sparse; margins nearly entire; costa almost smooth on abaxial surface; proximal laminal cells elongate, (19–) 42–54 (–84) × (5–) 7–9 (–12) µm; distal laminal cells irregularly rounded, elliptic or short-rectangular, (7–) 9–19 (–36) × (4–) 8–9 (–13) µm. Seta 2.5–3.5 cm.


Phenology: Capsules mature summer.
Habitat: Moist humus banks, humus on coniferous forest floors, and bogs on small conifers (often black spruce)
Elevation: 900-1300 m

Distribution

V27 585-distribution-map.gif

B.C., N.W.T., Que., Yukon, Europe

Discussion

Variety flexicaule is an arctic-alpine taxon that needs further collecting and study to determine its taxonomic status. J. Kucyniak (in D. Löve et al. 1958) called attention to it in the Quebec bryoflora and many European bryologists recognize it as a form (W. Mönkemeyer 1927), variety (J. Podpera 1954; L. I. Savicz-Lubitzkaya and Z. N. Smirnova 1970; V. M. Melnichuk 1970), or species (E. Nyholm 1986+, fasc. 1). It is immediately noticed because of the large size of the plants, whose stems are up to 18 cm, sparsely foliated with strongly falcate-secund leaves, and the long seta, 2.5–3.5 cm. It is further recognized microscopically by the irregular shape of the distal leaf cells, the nearly entire leaf margins, and the costa almost smooth on the abaxial surface.

Selected References

None.

Lower Taxa

None.
persistent +  and deciduous +
differentiated +
flexuose +
acute;obtuse +
Robert R. Ireland Jr. +
(Bridel) Wilson +
short to long +
differentiated +
narrowed +  and lanceolate +
Dicranum flexicaule +
microphyllous +
pseudomonoicous +, dioicous +, autoicous +  and sexual +
flagelliform +  and terete +
reproduction +  and asexual +
cucullate +
dark-brown +  and reddish-brown +
arcuate +  and straight +
inclined +  and horizontal +
0.1 cm1 mm <br />0.001 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
generally orange +  and brown +
specialized +
papillose +  and mammillose +
pseudomonoicous +, dioicous +  and sexual +
inflated +
differentiated +
1/6 +  and 1/4 +
papillose +
dioicous +  and sexual +
spreading +, erect-patent +  and erect-appressed +
smooth +  and rugose +
B.C. +, N.W.T. +, Que. +, Yukon +  and Europe +
reddish-brown +  and pitted-striolate +
900-1300 m +
Moist humus banks, humus on coniferous forest floors, and bogs on small conifers (often black spruce) +
2-stratose +  and 1-stratose +
enlarged +
falcate-secund +
sometimes abaxially +  and smooth +
smooth +  and slightly papillose +
acuminate +, ovate +  and lanceolate +
serrulate to strongly +
arcuate +  and straight +
long-rostrate +
0.1 cm1 mm <br />0.001 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
nonpitted +  and pitted +
toothed +  and forked +
short-acuminate +, concave +  and ovate +
pitted-striolate +
16 +  and single +
papillose +  and striolate +
Capsules mature summer. +
undifferentiated +
0.7 cm7 mm <br />0.007 m <br /> (1 cm10 mm <br />0.01 m <br />) +
0.4 cm4 mm <br />0.004 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
0.06 cm0.6 mm <br />6.0e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
Bryol. Brit., +
reddish-brown +  and white +
smooth +  and papillose +
1 +  and 2 +
well-developed +
brown;reddish yellow;yellow +
flexuose +  and straight +
0 +  and 6 +
elongate +
2.5 cm25 mm <br />0.025 m <br /> (3.5 cm35 mm <br />0.035 m <br />) +
1/3 +  and 1/2 +
spherical +
6 cm60 mm <br />0.06 m <br /> (18 cm180 mm <br />0.18 m <br />) +
8 cm80 mm <br />0.08 m <br /> (18 cm180 mm <br />0.18 m <br />) +
superficial +
Dicranum fuscescens var. flexicaule +
Dicranum fuscescens +
variety +
deciduous +
lanceolate +
green +, light green +  and dark brownish +