Didymodon anserinocapitatus

(X. J. Li) R. H. Zander

Bull. Buffalo Soc. Nat. Sci. 32: 162. 1993,.

Basionym: Barbula anserinocapitata X. J. Li Acta Bot. Yunnan. 3: 103, plate 2, figs. 1–9. 1981
Treatment appears in FNA Volume 27. Treatment on page 546. Mentioned on page 540, 547.

Plants green to reddish green. Stems to 1.5 cm, central strand present. Stem-leaves erect-appressed when dry, spreading and not keeled when moist, monomorphic, lanceolate, adaxially weakly concave across leaf, 0.7–1.1 mm (absent tip) to 2 mm whole, base scarcely differentiated in shape, margins recurved at mid leaf, entire, apex thickened, long-cylindric to clavate, usually soon deciduous, usually absent in mature leaves; costa excurrent, excurrency absent in mature leaves, not much widened or tapering at mid leaf but swollen in excurrency, pad of cells absent, adaxial costal cells quadrate, 4–6 cells wide at mid leaf, guide cells in 2 layers; basal laminal cells differentiated medially, walls thin; distal laminal cells 8–10 µm wide, 1: 1, papillae essentially absent, lumens angular, walls thin, weakly convex on both sides, 1-stratose except in deciduous apex. Specialized asexual reproduction by the deciduous leaf apex. Sexual condition and sporophytes unknown. Distal lamina KOH color reaction reddish orange.


Habitat: Red sandstone cliff, near river
Elevation: moderate to high elevations (1500-2200 m)

Distribution

V27 782-distribution-map.gif

Colo., N.Mex., Asia (w China), Asia (Kazakhstan), Asia (Siberia)

Discussion

In the flora area, Didymodon anserinocapitatus is known only from Colorado and New Mexico (R. H. Zander and W. A. Weber 1997). The arctic species D. johansenii is similar in the swollen, deciduous apex, but differs mainly by its distal laminal cells 13–15 µm wide, and guide cells in only a single layer. Didymodon rigidulus var. icmadophilus, a widespread montane taxon, is similar in appearance but its leaf apices are never swollen though sometimes fragile. The American material differs from the Asian type (China: Tibet, Nan Xian, Zang Mu 1704, isotype–NY) in being less robust, the leaves reaching only 1.8 mm, and the proximal cells quadrate to short-rectangular (leaves to 2 mm and proximal cells to 4:1 in the Asian specimen).

Selected References

None.

Lower Taxa

None.

"um" is not declared as a valid unit of measurement for this property."um" is not declared as a valid unit of measurement for this property.

quadrate +  and elongate +
0.3 cm3 mm <br />0.003 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
channeled +, concave +  and long-triangular +
0.04 cm0.4 mm <br />4.0e-4 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
of 1-2 rows +  and differentiated +
long-excurrent +  and short-excurrent +
deciduous +
Richard H. Zander +
(X. J. Li) R. H. Zander +
weak +  and strong +
ovate +  and crescent--shaped +
papillose +  and smooth +
usually slightly wider +
differentiated +
differentiated +
Barbula anserinocapitata +
obovoid +
cucullate +
stegocarpous +
monoicous +, dioicous +  and sexual +
flat to bulging +
thin;thick +
1 (?) +  and 10 (?) +
hexagonal +
tomentose +  and bare +
quadrate +  and rectangular +
occasionally weakly dentate or crenulate +  and entire +
subquadrate;hexagonal or rounded-angular occasionally shortly rectangular or rhomboid +
Colo. +, N.Mex. +, Asia (w China) +, Asia (Kazakhstan) +  and Asia (Siberia) +
moderate to high elevations (1500-2200 m) +
differentiated +
multicellular +
occasional +
Red sandstone cliff, near river +
yellow +  and orange-red or red +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.35 cm3.5 mm <br />0.0035 m <br />) +
concave;keeled;channeled;ovoid;lanceolate or lingulate +
ovate +, keeled +  and curled +
reduced +
specialized +
dioicous +  and sexual +
occasionally trigonous +  and weakly strongly convex +
short-conic;long-conic +
photosynthetic +
solid +  and simple +
lateral +  and terminal +
lateral +  and terminal +
twisted +, cleft +  and perforated +
red +, orange +  and yellow +
oblong +  and linear or long-triangular +
1 (?) +  and 2 (?) +
Bull. Buffalo Soc. Nat. Sci. +
straight +
yellowish;reddish-brown +
elongate +
0.3 cm3 mm <br />0.003 m <br /> (2 cm20 mm <br />0.02 m <br />) +
2 +  and 1 +
yellow +, red-orange +, red +  and laminal +
concave +, lanceolate +  and keeled +
2 cm20 mm <br />0.02 m <br /> (0.07 cm0.7 mm <br />7.0e-4 m <br />) +
rounded-triangular +  and rounded-pentagonal +
0 cm0 mm <br />0 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
Barbula sect. Acutae +, Barbula subsect. Acutiformes +, Barbula sect. Asteriscium +, Barbula sect. Fallaces +, Barbula subsect. Fallaciformes +, Barbula sect. Graciles +, Barbula sect. Rubiginosae +, Barbula sect. Vineales +, Barbula subsect. Vinealiformes +, Dactylhymenium +, Didymodon sect. Asteriscium +, Didymodon sect. Fallaces +, Didymodon sect. Graciles +, Didymodon sect. Vineales +, Geheebia +, Husnotiella +  and Trichostomopsis +
Didymodon anserinocapitatus +
Didymodon +
species +
elliptic;cylindric +
0.1 cm1 mm <br />0.001 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
rudimentary +
red +  and yellow +
specialized +
stemless +  and perigoniate +
green;reddish green +