Didymodon ferrugineus

(Schimper ex Bescherelle) M. O. Hill

J. Bryol. 11: 599. 1982,.

Basionym: Barbula ferruginea Schimper ex Bescherelle Mém. Soc. Natl. Sci. Nat. Cherbourg 16: 181. 1872
Synonyms: Barbula reflexa (Bridel) Bridel Didymodon fallax var. reflexus (Bridel) R. H. Zander Didymodon rigidicaulis
Treatment appears in FNA Volume 27. Treatment on page 557. Mentioned on page 541, 560, 581.

Plants usually redbrown. Stems to 2.5 cm, central strand present. Stem-leaves appressed to spreading when dry, strongly recurved and keeled when moist, monomorphic, ovate-triangular to ovatelanceolate, adaxially grooved along costa, 0.8–2 mm, base scarcely differentiated in shape to ovate, margins nearly plane to recurved at mid leaf, usually entire, apex broadly acute, not fragile; costa percurrent to short-excurrent, weakly tapering and not much wider at the base, pad of cells absent, adaxial costal cells elongate, 2–4 cells wide at mid leaf, guide cells in 1 layer; basal laminal cells little differentiated, quadrate or very shortrectangular, walls thickened; distal laminal cells 11–15 µm wide, 1: 1, papillae usually present, simple, often large and sharp, 1–3 centered over lumens, lumens angular, walls irregularly thickened, convex on both sides, 1-stratose. Specialized asexual reproduction specialized structures absent. Seta 0.6–1.2 cm. Capsule 0.7–1.5 mm; peristome teeth 16, linear, cleft to near base, nearly straight to twisted counterclockwise once, 275–600 µm. Spores 7–9 µm. Distal laminal KOH reaction red or orangebrown.


Phenology: Capsules mature winter–spring.
Habitat: Soil, ledges and outcrops, limestone, tundra, wet areas
Elevation: low to moderate elevations (60-1400 m)

Distribution

V27 800-distribution-map.gif

Greenland, Alta., B.C., Man., N.B., Nfld. and Labr., N.W.T., Nunavut, Ont., Yukon, Alaska, Ariz., Ark., Colo., Ill., Mich., Mont., N.Mex., N.Y., N.C., Vt., Va., W.Va., Mexico, West Indies, Central America, Europe, Asia, n Africa

Discussion

The sporophyte of Didymodon ferrugineus is rare and similar to, but often smaller than, that of D. fallax. This species is easily mistaken for the western Triquetrella californica, which, however, is quickly distinguished by its triangular stem section, distal leaf margins sharply crenulate by projecting papillae (but not by projecting cell walls as in D. nigrescens), and distal medial laminal papillae tall, branching from the base, and centered over each lumen. It intergrades with D. fallax to some extent but is usually larger, with broader, ovate-lanceolate leaves that are commonly strongly reflexed (and therefore do not lie flat on a slide), and the papillae may be large and strongly evident. The leaf base is never abruptly ovate as it sometimes is in D. fallax. The peristome is very fragile and often appears missing.

The illegitimate name Tortula recurvifolia (Schimper) Austin has also been used for this species.

Selected References

None.

Lower Taxa

None.

"um" is not declared as a valid unit of measurement for this property."um" is not declared as a valid unit of measurement for this property.

... more about "Didymodon ferrugineus"
quadrate +  and elongate +
0.3 cm3 mm <br />0.003 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
channeled +, concave +  and long-triangular +
0.04 cm0.4 mm <br />4.0e-4 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
of 1-2 rows +  and differentiated +
long-excurrent +  and short-excurrent +
not fragile +
Richard H. Zander +
(Schimper ex Bescherelle) M. O. Hill +
weak +  and strong +
ovate +  and crescent--shaped +
papillose +  and smooth +
usually slightly wider +
ovate;differentiated +
differentiated +
Barbula ferruginea +
obovoid +
cucullate +
stegocarpous +
0.07 cm0.7 mm <br />7.0e-4 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
monoicous +, dioicous +  and sexual +
flat to bulging +
thin;thick +
1 (?) +  and 10 (?) +
hexagonal +
tomentose +  and bare +
percurrent +  and short-excurrent +
tapering +
quadrate +  and rectangular +
occasionally weakly dentate or crenulate +  and entire +
subquadrate;hexagonal or rounded-angular occasionally shortly rectangular or rhomboid +
Greenland +, Alta. +, B.C. +, Man. +, N.B. +, Nfld. and Labr. +, N.W.T. +, Nunavut +, Ont. +, Yukon +, Alaska +, Ariz. +, Ark. +, Colo. +, Ill. +, Mich. +, Mont. +, N.Mex. +, N.Y. +, N.C. +, Vt. +, Va. +, W.Va. +, Mexico +, West Indies +, Central America +, Europe +, Asia +  and n Africa +
low to moderate elevations (60-1400 m) +
differentiated +
multicellular +
occasional +
Soil, ledges and outcrops, limestone, tundra, wet areas +
yellow +  and orange-red or red +
grooved +  and monomorphic +
0.08 cm0.8 mm <br />8.0e-4 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
recurved +, appressed +  and spreading +
ovate-triangular;ovatelanceolate +
ovate +, keeled +  and curled +
reduced +
specialized +
dioicous +  and sexual +
occasionally trigonous +  and weakly strongly convex +
short-conic;long-conic +
photosynthetic +
1 +  and 3 +
lateral +  and terminal +
lateral +  and terminal +
red +, orange +  and yellow +
oblong +  and linear or long-triangular +
Capsules mature winter–spring. +
1 (?) +  and 2 (?) +
straight +
yellowish;reddish-brown +
elongate +
0.6 cm6 mm <br />0.006 m <br /> (1.2 cm12 mm <br />0.012 m <br />) +
2 +  and 1 +
orangebrown +  and red +
rounded-triangular +  and rounded-pentagonal +
0 cm0 mm <br />0 m <br /> (2.5 cm25 mm <br />0.025 m <br />) +
absent +  and asexual +
Barbula reflexa +, Didymodon fallax var. reflexus +  and Didymodon rigidicaulis +
Didymodon ferrugineus +
Didymodon +
species +
elliptic;cylindric +
0.1 cm1 mm <br />0.001 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
rudimentary +
red +  and yellow +
stemless +  and perigoniate +