familyPottiaceae
subfamilyPottiaceae subfam. Barbuloideae
genusDidymodon
speciesDidymodon rigidulus
varietyDidymodon rigidulus var. subulatus

Didymodon rigidulus var. subulatus

(Thériot & E. B. Bartram) R. H. Zander

Cryptog. Bryol. Lichénol. 2: 395. 1981,.

Richard H. Zander
Basionym: Didymodon mexicanus var. subulatus Thériot & E. B. Bartram Bryologist 29: 1, plate 1. 1926
Treatment appears in FNA Volume 27. Treatment on page 544. Mentioned on page 543, 561.
Please click on the illustration for a higher resolution version.
Illustrator: Patricia M. Eckel

Copyright: Flora of North America Association

Stem-leaves appressed to weakly spreading when dry, longlanceolate, evenly broadened, long-oval; costa excurrent as a subula; proximal cells rectangular; distal laminal cells smooth, with quadrate lumens, lamina 2-stratose in distal 1/2–3/4; gemmae absent. Peristome teeth short, rudimentary, of 16 oval teeth with 1–2 (–4) articulations.


Phenology: Capsule maturity unknown.
Habitat: Rocks, ledges, soil, canyons
Elevation: moderate to high elevations (1000-2000 m)

Distribution

V27 777-distribution-map.gif

Ariz., Tex., Mexico (Distrito Federal), Mexico (Michoacán), Mexico (Puebla), Mexico (Querétaro), Mexico (Sonora)

Discussion

Variety subulatus is similar to var. icmadophilus in leaf shape and most other characters, and intergrades occur. It may be regarded as a somewhat distinctive geographical variant. The variety phenocopies Grimmia hamulosa, which differs by the grimmiaceous peristome, homogeneous costal section, and more robust habit.

Selected References

None.

Lower Taxa

None.

"um" is not declared as a valid unit of measurement for this property."um" is not declared as a valid unit of measurement for this property.

... more about "Didymodon rigidulus var. subulatus"
RDF feed
Abaxial epidermis fragility
weak +
Abaxial epidermis presence
absent +
Abaxial epidermis variability
differentiated +
Adaxial cell architecture
short-excurrent +
Adaxial cell shape
quadrate +  and elongate +
Adaxial epidermis presence
absent +
Adaxial pad pubescence or shape
bulging +
Adaxial pad quantity
lacking +
Adaxial surface atypical some measurement
0.3 cm3 mm <br />0.003 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
Adaxial surface shape
channeled +, concave +  and long-triangular +
Adaxial surface some measurement
0.04 cm0.4 mm <br />4.0e-4 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
Annulus architecture
revoluble +
Annulus duration
deciduous +
Annulus shape or vernation
revolute +
Annulus variability
of 1-2 rows +  and differentiated +
Apex architecture
2-stratose +
Apex duration
caducous +
Apex fragility
fragile +
Apex shape
acute +  and subulate +
Articulation atypical quantity
2 (4) +
Articulation quantity
1 +  and 2 +
Author
Richard H. Zander +
Authority
(Thériot & E. B. Bartram) R. H. Zander +
Axillary hair quantity
5 +
Band fragility
weak +  and strong +
Band presence
absent +
Band shape
ovate +  and crescent--shaped +
Basal laminal cell pubescence or shape
bulging +
Basal laminal cell quantity
1 (?) +
Basal laminal cell relief
papillose +  and smooth +
Basal laminal cell shape
Rectangular (?) +
Basal laminal cell variability
differentiated +
Basal laminal cell width
usually slightly wider +
Basal membrane architecture
trabeculate +
Basal membrane height
high +
Basal membrane position
low +
Basal membrane presence
absent +
Base shape
ovate +
Base variability
differentiated +
Basionyms
Didymodon mexicanus var. subulatus +
Body shape
obovoid +
Branchlet height or length or size
short +
Calyptra architecture or pubescence or relief
smooth +
Calyptra relief
papillose +
Calyptra shape
cucullate +
Capsule dehiscence
stegocarpous +
Capsule some measurement
0.1 cm1 mm <br />0.001 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
Cell-wall development
specialized +
Cell-wall prominence or shape
flat +
Cell-wall pubescence or shape
bulging +
Cell-wall reproduction
monoicous +, dioicous +  and sexual +
Cell-wall shape
flat to bulging +
Cell-wall size or width
thickened +
Cell-wall width
thin;thick +
Cell architecture
vesiculose +
Cell coloration
brownish +
Cell length or size
long +
Cell quantity
1 +  and 10 +
Cell shape
hexagonal +
Cell variability
differentiated +
Central scleroderm presence
absent +
Central strand fusion
distinct +
Central strand presence
absent +
Central strand pubescence
tomentose +  and bare +
Costa architecture
excurrent +
Costa architecture or shape
spurred +
Costa shape
tapering +  and channeled +
Costa width
widened +
Costal adaxial cell shape
quadrate +
Distal cell position
higher +
Distal cell pubescence or shape
bulging +
Distal cell quantity
2 (4) +
Distal cell shape
quadrate +  and rectangular +
Distal cell width
Wider (Wider) +
Distal laminal cell architecture
2-stratose +
Distal laminal cell architecture or pubescence or relief
smooth +
Distal laminal cell arrangement
stacked +
Distal laminal cell quantity
1 (?) +
Distal laminal cell shape
Rhomboid (?) +, Rarely (?) +, Hexagonal (?) +  and Subquadrate (?) +
Distal laminal cell size
small +
Distal laminal cell width
8um;11um +
Distal margin architecture
2-stratose +
Distal margin architecture or shape
occasionally weakly dentate or crenulate +  and entire +
Distal margin orientation
recurved +
Distal margin shape
plane +
Distal margin shape or vernation
revolute +
Distal medial cell architecture
1-stratose +
Distal medial cell quantity
1 +
Distal medial cell shape
subquadrate;hexagonal or rounded-angular occasionally shortly rectangular or rhomboid +
Distal medial cell width
8um (13um) +
Distribution
Ariz. +, Tex. +, Mexico (Distrito Federal) +, Mexico (Michoacán) +, Mexico (Puebla) +, Mexico (Querétaro) +  and Mexico (Sonora) +
Division architecture
spiculose +
Division shape
filiform +
Elevation
moderate to high elevations (1000-2000 m) +
Epidermis variability
differentiated +
Gemma architecture
multicellular +
Gemma position
axillary +
Gemma presence
absent +
Gemma quantity
occasional +
Gemma reproduction
asexual +
Gemma shape
Ovate (?) +  and Elliptic (?) +
Guide cell quantity
2 +  and 6 +
Guide cell shape or vernation
revolutus +
Guide cell size
small +
Habitat
Rocks, ledges, soil, canyons +
Hyalodermi presence
absent +
Hyalodermi quantity
rare +
Hydroid strand presence
absent +
Illustration
Present +
Illustration copyright
Flora of North America Association +
Illustrator
Patricia M. Eckel +
Interior leaf architecture or shape
sheathing +
Interior leaf shape
ovate +
Interior leaf size
enlarged +
Koh coloration
yellow +  and orange-red or red +
Koh quantity
2% +
Lamina architecture
2-stratose +
Laminal cell shape
Rhombiodal-rectangular (?) +
Layer atypical quantity
1 (2) +
Layer quantity
single +
Leaf-stem architecture
long-ligulate +  and monomorphic +
Leaf-stem arrangement or shape
contorted +
Leaf-stem atypical distance
0.17 cm1.7 mm <br />0.0017 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
Leaf-stem distance
0.08 cm0.8 mm <br />8.0e-4 m <br /> (0.17 cm1.7 mm <br />0.0017 m <br />) +
Leaf-stem fixation or orientation
appressed +
Leaf-stem orientation
not +, spreading +, appressed-incurved +  and weakly spreading +
Leaf-stem shape
channeled;ovatelanceolate;keeled +
Leaf architecture
twisted +
Leaf arrangement
crowded +
Leaf fragility
fragile +
Leaf orientation
spreading +  and strongly reflexed +
Leaf shape
ovate +, keeled +  and curled +
Leaf size
reduced +
Lumen development
specialized +
Lumen reproduction
dioicous +  and sexual +
Lumen shape
quadrate +
Margin architecture
bordered +
Margin architecture or shape
entire +
Margin orientation
recurved +
Margin shape
plane +
Margin shape or vernation
involute +
Neck presence
absent +
Neck size
small +
Number of lower taxa
0 +
Operculum height or length or size
short +
Operculum shape
short-conic;long-conic +
Outgrowth nutrition
photosynthetic +
Pair presence
absent +
Papilla architecture
simple +
Papilla presence
absent +
Papilla quantity
multiplex +
Papilla shape
2-fid +
Perichaetial leaf architecture or shape
sheathing +
Perichaetial leaf shape
long-oval +
Perichaetial leaf size
usually larger +
Perichaetium position
lateral +  and terminal +
Perichaetium position or structure subtype
terminal +
Perigonium position
lateral +  and terminal +
Peristome presence
absent +
Peristome tooth architecture
twisted +, cleft +  and perforated +
Peristome tooth arrangement
grouped +
Peristome tooth arrangement or course or shape
linear +
Peristome tooth coloration
red-orange +  and yellow +
Peristome tooth course
straight +
Peristome tooth height or length or size
short +
Peristome tooth orientation
erect +
Peristome tooth position or shape
distal +
Peristome tooth presence
absent +
Peristome tooth prominence
rudimentary +
Peristome tooth quantity
0-740 um +  and occasionally absent +
Peristome tooth shape
0-740 um-triangular +  and long-triangular +
Peristome tooth structure in adjective form
laminal +
Phenology
Capsule maturity unknown. +
Plant size
small +
Projection presence
absent +
Projection quantity
3 (5) +
Proximal cell quantity
1 (?) +  and 2 (?) +
Proximal cell shape
Rectangular (?) +
Proximal margin shape
decurrent +
Proximal strand quantity
1 (3) +
Publication title
Cryptog. Bryol. Lichénol. +
Reference
None +
Row atypical quantity
4 (5) +
Row course
straight +
Row orientation or shape
oblique +
Row quantity
3;4 +
Scleroderm presence
absent +
Seta architecture
twisted +
Seta architecture or arrangement or growth form
solitary +
Seta coloration
yellowish;reddish-brown +
Seta shape
elongate +
Seta some measurement
0.7 cm7 mm <br />0.007 m <br /> (1.7 cm17 mm <br />0.017 m <br />) +
Side quantity
2 +  and 1 +
Source xml
https://bibilujan@bitbucket.org/aafc-mbb/fna-fine-grained-xml.git/src/84153f6d59a0a91d69695978a64cee7560374f8e/V27/V27 777.xml +
Spore architecture
twisted +
Spore coloration
yellow +, red-orange +, red +  and laminal +
Spore size
larger +
Spore some measurement
9um (12um) +
Stem-leaf orientation
appressed;weakly spreading +
Stem-leaf shape
long-oval +
Stem-leaf width
broadened +
Stem architecture
branching +
Stem architecture or shape
half-sheathing +
Stem atypical some measurement
2 cm20 mm <br />0.02 m <br /> (9 cm90 mm <br />0.09 m <br />) +
Stem presence
absent +
Stem shape
rounded-triangular +  and rounded-pentagonal +
Stem some measurement
0 cm0 mm <br />0 m <br /> (2 cm20 mm <br />0.02 m <br />) +
Subulum length or size
long +
Subulum shape
blunt +
Subulum width
thick +
Synonyms
Barbula rigidula +
Taxon family
Pottiaceae +
Taxon name
Didymodon rigidulus var. subulatus +
Taxon parent
Didymodon rigidulus +
Taxon rank
variety +
Theca shape
elliptic;cylindric +
Theca some measurement
0.1 cm1 mm <br />0.001 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
Tooth prominence
rudimentary +
Tooth quantity
16 (?) +
Tooth shape
oval +
Transverse-section shape
Reniform (Semicircular) +
Tuber size
small +
Volume
Volume 27 +
Wall architecture
porose +
Wall coloration
red +  and yellow +
Wall development
specialized +
Wall pubescence or shape
bulging +
Wall relief
Smooth (?) +  and Papillose (?) +
Wall size or width
thickened +
Wall width
thin +  and evenly thickened +
Whole organism architecture
stemless +  and perigoniate +
Whole organism architecture or shape
budlike +
Whole organism coloration
blackish;green +
Whole organism growth form
plant +
Whole organism size
small +