Didymodon vinealis var. rubiginosus

(Mitten) R. H. Zander

Cryptog. Bryol. Lichénol. 2: 417. 1981,.

Basionym: Barbula rubiginosa Kindberg J. Linn. Soc., Bot. 8: 27, fig. 5. 1865
Synonyms: Barbula melanocarpa Kindberg Barbula subicmadophila R. H. Zander Didymodon occidentalis Cardot & Thériot Racomitrium cyclodictyon
Treatment appears in FNA Volume 27. Treatment on page 552. Mentioned on page 551.

Capsule with peristome absent; operculum narrowly conic, cells straight or nearly so.


Phenology: Capsules mature spring.
Habitat: Moderate elevations

Distribution

V27 792-distribution-map.gif

B.C., Colo., Idaho, Mont., Oreg., Wash.

Discussion

Although the gametophyte of var. rubiginosus is little variable, having short-lanceolate to triangular leaves, it is well within the range of variation of the typical variety. It has been said to differ in a very narrow distal lamina that is often fragile or notched, or by large distal laminal cells, or by quadrate proximal cells, but these characters are insufficient to distinguish sterile specimens. The sporophyte is required for accurate identification, with the absence (not mere fragility) of the peristome as the critical character, following the species concept of W. C. Steere (1938b); the nearly straight opercular cells are diagnostic. Forms of var. vinealis with weakly twisted opercular cells and weak, very fragile peristomes (e. g. California, Flowers 6561, COLO) may be taken as intermediates. B. H. Allen (1995) recently provided new American synonymy. The Mexican species Didymodon incrassatolimbatus Cardot should be looked for in the flora area; it differs by the broadly lanceolate, marginally 2-stratose leaves and well developed though short, red, erect peristome.

Selected References

None.

Lower Taxa

None.

"um" is not declared as a valid unit of measurement for this property."um" is not declared as a valid unit of measurement for this property.

quadrate +  and elongate +
0.3 cm3 mm <br />0.003 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
channeled +, concave +  and long-triangular +
0.04 cm0.4 mm <br />4.0e-4 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
of 1-2 rows +  and differentiated +
long-excurrent +  and short-excurrent +
not fragile +
Richard H. Zander +
(Mitten) R. H. Zander +
weak +  and strong +
ovate +  and crescent--shaped +
papillose +  and smooth +
across the leaf +  and differentiated +
usually slightly wider +
differentiated +
Barbula rubiginosa +
obovoid +
cucullate +
stegocarpous +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.25 cm2.5 mm <br />0.0025 m <br />) +
monoicous +, dioicous +  and sexual +
flat to bulging +
thin;thick +
nearly +  and straight +
1 (?) +  and 10 (?) +
conical +
tomentose +  and bare +
short-excurrent +  and percurrent +
tapering +
widened +
quadrate +  and rectangular +
occasionally weakly dentate or crenulate +  and entire +
subquadrate;hexagonal or rounded-angular occasionally shortly rectangular or rhomboid +
B.C. +, Colo. +, Idaho +, Mont. +, Oreg. +  and Wash. +
differentiated +
multicellular +
Moderate elevations +
yellow +  and orange-red or red +
grooved +, monomorphic +  and twisted +
0.25 cm2.5 mm <br />0.0025 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
spreading-flexuose +
0.08 cm0.8 mm <br />8.0e-4 m <br /> (0.25 cm2.5 mm <br />0.0025 m <br />) +
spreading +  and spreading-recurved +
lanceolate;keeled +
ovate +, keeled +  and curled +
reduced +
oval to roundedquadrate +  and irregular +
specialized +
dioicous +  and sexual +
occasionally trigonous +  and weakly strongly convex +
photosynthetic +
2 +  and 3 +
lateral +  and terminal +
lateral +  and terminal +
red +, orange +  and yellow +
oblong +  and linear or long-triangular +
Capsules mature spring. +
1 (?) +  and 2 (?) +
Cryptog. Bryol. Lichénol. +
straight +
yellowish;reddish-brown +
elongate +
0.8 cm8 mm <br />0.008 m <br /> (1 cm10 mm <br />0.01 m <br />) +
2 +  and 1 +
red-orange +, light +  and dark redbrown +
rounded-triangular +  and rounded-pentagonal +
0 cm0 mm <br />0 m <br /> (2 cm20 mm <br />0.02 m <br />) +
Barbula melanocarpa +, Barbula subicmadophila +, Didymodon occidentalis +  and Racomitrium cyclodictyon +
Didymodon vinealis var. rubiginosus +
Didymodon vinealis +
variety +
elliptic;cylindric +
0.1 cm1 mm <br />0.001 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
rudimentary +
2-stratose +, 1-stratose +  and not perforated +
red +  and yellow +
specialized +
convex +, quadrate +  and rectangular +
thickened +, thin +  and weakly thickened +
stemless +  and perigoniate +