Didymodon vinealis var. vinealis

Synonyms: Barbula bakeri Cardot & Thériot Barbula circinnulata Müller Hal. & Kindberg Barbula cylindrica (Taylor) Schimper Barbula flexifolia Hampe Barbula horridifolia Müller Hal. & Kindberg Barbula laterita Kindberg Barbula pseudorigidula Müller Hal. & Kindberg Barbula robustifolia Müller Hal. & Kindberg Barbula semitorta Sullivant Barbula subcylindrica Brotherus Barbula subgracilis Müller Hal. & Kindberg Barbula tortellifolia Müller Hal. & Kindberg Barbula treleasei Cardot & Thériot Barbula vinealis var. flaccida (Bruch & Schimper) R. H. Zander Barbula virescens Didymodon vinealis var. flaccidus
Treatment appears in FNA Volume 27. Treatment on page 551. Mentioned on page 552.

Capsule with peristome present, twisted, to 1300 µm; operculum broadly conic, cells twisted.


Phenology: Capsules mature spring–summer.
Habitat: Soil, calcareous rock, granite outcrop, schist, sandstone
Elevation: moderate to high elevations (600-2300 m)

Distribution

V27 791-distribution-map.gif

Alta., B.C., Nunavut, Alaska, Ariz., Calif., Colo., Idaho, Ill., Mont., Nev., N.Mex., Oreg., Utah, Wash., Wyo., Mexico, Central America, South America, Europe, Asia (Middle East), Asia (Siberia), n Africa, Atlantic Islands (Iceland), Pacific Islands

Discussion

In Nunavut, var. vinealis is known from Banks Island.

Some collections of var. vinealis have elongate, very fragile apices, sometimes 2-stratose. Variety flaccidus was synonymized by P. Sollman (1983), but may be distinguished if needed by the leaves long, often 2.5 mm or longer, and crisped when dry, the distal margins plane. The peristome commonly falls with the operculum in the specimens seen, and may appear to be absent, but the operculum has twisted cells and is thin-walled.

The leaves of Didymodon vinealis var. vinealis may be nearly plane margined and quite fragile (Colorado: Jefferson County, Weber and Wittman B-110966, MO, COLO), with the appearance of variants of Trichostomum tenuirostre, but the distal laminal cells are distinctly reddish in KOH under high magnification (not yellow).

Selected References

None.

Lower Taxa

None.

"um" is not declared as a valid unit of measurement for this property."um" is not declared as a valid unit of measurement for this property."um" is not declared as a valid unit of measurement for this property.

quadrate +  and elongate +
0.3 cm3 mm <br />0.003 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
channeled +, concave +  and long-triangular +
0.04 cm0.4 mm <br />4.0e-4 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
of 1-2 rows +  and differentiated +
long-excurrent +  and short-excurrent +
not fragile +
Richard H. Zander +
(Bridel) R. H. Zander +
weak +  and strong +
ovate +  and crescent--shaped +
papillose +  and smooth +
across the leaf +  and differentiated +
usually slightly wider +
differentiated +
Barbula vinealis +
obovoid +
cucullate +
stegocarpous +
monoicous +, dioicous +  and sexual +
flat to bulging +
thin;thick +
1 +  and 10 +
conical +
tomentose +  and bare +
short-excurrent +  and percurrent +
tapering +
widened +
quadrate +  and rectangular +
occasionally weakly dentate or crenulate +  and entire +
subquadrate;hexagonal or rounded-angular occasionally shortly rectangular or rhomboid +
Alta. +, B.C. +, Nunavut +, Alaska +, Ariz. +, Calif. +, Colo. +, Idaho +, Ill. +, Mont. +, Nev. +, N.Mex. +, Oreg. +, Utah +, Wash. +, Wyo. +, Mexico +, Central America +, South America +, Europe +, Asia (Middle East) +, Asia (Siberia) +, n Africa +, Atlantic Islands (Iceland) +  and Pacific Islands +
moderate to high elevations (600-2300 m) +
differentiated +
multicellular +
Soil, calcareous rock, granite outcrop, schist, sandstone +
yellow +  and orange-red or red +
grooved +, monomorphic +  and twisted +
0.25 cm2.5 mm <br />0.0025 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
spreading-flexuose +
0.08 cm0.8 mm <br />8.0e-4 m <br /> (0.25 cm2.5 mm <br />0.0025 m <br />) +
spreading +  and spreading-recurved +
lanceolate;keeled +
spreading +  and strongly reflexed +
ovate +, keeled +  and curled +
reduced +
oval to roundedquadrate +  and irregular +
specialized +
dioicous +  and sexual +
occasionally trigonous +  and weakly strongly convex +
photosynthetic +
2 +  and 3 +
lateral +  and terminal +
lateral +  and terminal +
red +, orange +  and yellow +
oblong +  and linear or long-triangular +
Capsules mature spring–summer. +
1 (?) +  and 2 (?) +
straight +
yellowish;reddish-brown +
elongate +
0.8 cm8 mm <br />0.008 m <br /> (1 cm10 mm <br />0.01 m <br />) +
2 +  and 1 +
red-orange +, light +  and dark redbrown +
2 cm20 mm <br />0.02 m <br /> (9 cm90 mm <br />0.09 m <br />) +
rounded-triangular +  and rounded-pentagonal +
0 cm0 mm <br />0 m <br /> (2 cm20 mm <br />0.02 m <br />) +
Barbula bakeri +, Barbula circinnulata +, Barbula cylindrica +, Barbula flexifolia +, Barbula horridifolia +, Barbula laterita +, Barbula pseudorigidula +, Barbula robustifolia +, Barbula semitorta +, Barbula subcylindrica +, Barbula subgracilis +, Barbula tortellifolia +, Barbula treleasei +, Barbula vinealis var. flaccida +, Barbula virescens +  and Didymodon vinealis var. flaccidus +
Didymodon vinealis var. vinealis +
Didymodon vinealis +
variety +
elliptic;cylindric +
0.1 cm1 mm <br />0.001 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
rudimentary +
2-stratose +, 1-stratose +  and not perforated +
red +  and yellow +
specialized +
convex +, quadrate +  and rectangular +
thickened +, thin +  and weakly thickened +
stemless +  and perigoniate +