Diphasiastrum tristachyum

(Pursh) Holub

Preslia 47: 108. 1975.

Common names: Blue ground-cedar lycopode à trois épis
Basionym: Lycopodium tristachyum Pursh Fl. Amer. Sept. 2: 653. 1814
Treatment appears in FNA Volume 2.

Horizontal stems deeply (5–12 cm) buried, 1.5–3.2 mm wide; leaves spatulate to somewhat obovate, 1.8–3.5 × 1.1–1.5, apex faintly erose to irregularly lobed. Upright shoots clustered, branching near base, 17–36 cm; leaves monomorphic, appressed, subulate, 1.9–3.4 × 0.6–1 mm, apex acute. Branchlets square with rounded angles in cross-section, 1–2.2 mm wide, annual bud constrictions abrupt and conspicuous; upperside convex, bluish to whitish green. Leaves on branchlets 4-ranked, upperside leaves appressed, needlelike, free portion of blade 1–1.7 × 0.5–0.9 mm; lateral leaves appressed, 3.4–7.2 × 1.1–2 mm; underside leaves appressed, somewhat decurrent, 1–2 × 0.4–0.7 mm. Peduncles (1–) 3, 4–15 × 0.4–1 mm; leaves remote, scattered, decurrent, free tips ascending, subulate, 2–3 × 0.2–0.25 mm. Stalks mostly formed by successive forking of peduncle, branches uniformly spaced. Strobili (2–) 3–4 (–7), 10–28 × 2–3 mm, apex round-tipped, sterile tips absent. Sporophylls deltate, 2.2–3.5 × 1.6–3 mm, apex gradually tapering. 2n = 46.


Habitat: Sterile, acidic soils in open conifer and oak forests
Elevation: 50–1800 m

Distribution

V2 573-distribution-map.gif

Man., N.B., Nfld. and Labr. (Nfld.), N.S., Ont., P.E.I., Que., Ala., Conn., Del., D.C., Ga., Ind., Ky., Maine, Md., Mass., Mich., Minn., Mo., N.H., N.J., N.Y., N.C., Ohio, Pa., R.I., S.C., Tenn., Vt., Va., W.Va., Wis., Europe, Asia in w China

Discussion

The distinctive Diphasiastrum tristachyum has narrow, rounded branches and dull, bluish white color. It is a parent in more hybrid combinations than any other North American Diphasiastrum.

The reticulogram shows the known pattern of interspecific hybridization in Diphasiastrum. The hybrids are discussed in detail by J. H. Wilce (1965), and their cytology is summarized by F. S. Wagner (1992). The best known North American hybrids are the four involving D. tristachyum. All of the hybrids have apparently normal meiosis and spores.

Diphasiastrum × zeilleri (Rouy) Holub (= D. complanatum X tristachyum) is a frequent plant in areas of distributional overlap between the parents, especially in north central and western Minnesota jackpine forests.

Diphasiastrum × habereri (House) Holub (= D. digitatum X tristachyum) has been overlooked and confused with both parents in zones of overlap. It is found occasionally to frequently in habitats like those of the parents, not necessarily growing close to them.

Diphasiastrum × issleri (Rouy) Holub (= D. alpinum X tristachyum) is a rare hybrid in North America, reported only from Maine, but much more widespread in Europe.

Diphasiastrum × sabinifolium (Willdenow) Holub (= D. sitchense X tristachyum) is widespread and frequent in eastern Canada. This hybrid is commonly confused with D. sitchense. It is highly variable, and some individuals approach one or the other parent in morphology (W. J. Cody and D. M. Britton 1989). In the flora, the populations are mainly disjunct from the main range of D. sitchense, including those in Ontario, Quebec, Maine, Michigan, New Hampshire, New York, Pennsylvania, and Vermont.

Two other North American nothospecies are Diphasiastrum complanatum X digitatum and D. alpinum X sitchense.

Selected References

None.

Lower Taxa

None.
... more about "Diphasiastrum tristachyum"
round-tipped +
Warren H. Wagner Jr. +  and Joseph M. Beitel +
(Pursh) Holub +
Lycopodium tristachyum +
0.1 cm1 mm <br />0.001 m <br /> (0.22 cm2.2 mm <br />0.0022 m <br />) +
Blue ground-cedar +  and lycopode à trois épis +
Man. +, N.B. +, Nfld. and Labr. (Nfld.) +, N.S. +, Ont. +, P.E.I. +, Que. +, Ala. +, Conn. +, Del. +, D.C. +, Ga. +, Ind. +, Ky. +, Maine +, Md. +, Mass. +, Mich. +, Minn. +, Mo. +, N.H. +, N.J. +, N.Y. +, N.C. +, Ohio +, Pa. +, R.I. +, S.C. +, Tenn. +, Vt. +, Va. +, W.Va. +, Wis. +, Europe +  and Asia in w China +
50–1800 m +
photosynthetic +
carrot--shaped +
Sterile, acidic soils in open conifer and oak forests +
0.34 cm3.4 mm <br />0.0034 m <br /> (0.72 cm7.2 mm <br />0.0072 m <br />) +
1.1mm +  and 2mm +
pseudomonopodial +, dichotomous +, branching +, branched +  and simple +
flat +  and round +
monomorphic +
remote;dense +
1.9mm;3.4mm +
1.1 (?) +  and 1.5 (?) +
decurrent +, subulate +, obovate +  and spatulate to somewhat +
larger +
trimorphic +  and uniform +
0.06 cm0.6 mm <br />6.0e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
aerial +  and subterranean +
0.4 cm4 mm <br />0.004 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
0.04 cm0.4 mm <br />4.0e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
0.1 cm1 mm <br />0.001 m <br /> (0.17 cm1.7 mm <br />0.0017 m <br />) +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.09 cm0.9 mm <br />9.0e-4 m <br />) +
emerging +
0.2 cm2 mm <br />0.002 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
quadrate +  and flattened +
17 cm170 mm <br />0.17 m <br /> (36 cm360 mm <br />0.36 m <br />) +
plectostelic +  and actino +
axillary +  and adaxial +
thick-walled +  and trilete +
0.22 cm2.2 mm <br />0.0022 m <br /> (0.35 cm3.5 mm <br />0.0035 m <br />) +
1.6mm +  and 3mm +
protostelic +
1.5mm;3.2mm +
subterranean +
stalked +  and sessile +
modified +  and unmodified +
photosynthetic +
0.2 cm2 mm <br />0.002 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
small-grooved +
subterranean +
reticulate +, rugulate +  and pitted +
Diphasiastrum tristachyum +
Diphasiastrum +
species +
2mm;3mm +
subulate +
0.02 cm0.2 mm <br />2.0e-4 m <br /> (0.025 cm0.25 mm <br />2.5e-4 m <br />) +
0.1 cm1 mm <br />0.001 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
0.04 cm0.4 mm <br />4.0e-4 m <br /> (0.07 cm0.7 mm <br />7.0e-4 m <br />) +
bluish +  and whitish green +