Draba densifolia

Nuttall in J. Torrey and A. Gray

in J. Torrey and A. Gray, Fl. N. Amer. 1: 104. 1838.

IllustratedEndemic
Synonyms: Draba caeruleomontana Payson & H. St. John Draba caeruleomontana var. piperi Payson & H. St. John Draba glacialis var. pectinata S. Watson Draba globosa var. sphaerula (J. F. Macbride & Payson) O. E. Schulz Draba mulfordiae J. F. Macbride & Payson Draba nelsonii Hooker Draba oligosperma var. pectinata (S. Watson) Jepson Draba pectinata J. F. Macbride & Payson Draba sphaerula
Treatment appears in FNA Volume 7. Treatment on page 306. Mentioned on page 280, 282, 284, 288, 308.

Perennials; (cespitose, pulvinate); caudex branched (dense with persistent leaf remains, branches sometimes terminating in sterile rosettes); scapose. Stems unbranched, (0.05–) 0.2–1 (–1.7) dm, usually glabrous, rarely pubescent, trichomes usually simple, 0.3–0.8 mm, and 2–4 (or 5) -rayed, 0.1–0.6 mm, (rarely predominantly simple ones). Basal leaves rosulate; sessile; blade linear to oblong or oblanceolate-linear, 0.3–0.9 (–1.4) cm × 0.5–1.5 (–2) mm, margins entire, (ciliate, trichomes simple, 0.3–1.2 mm), surfaces glabrous or sparsely pubescent, abaxially with short-stalked, 2–4-rayed trichomes, 0.1–0.3 mm (midvein prominent), adaxially rarely with subapical, simple trichomes. Cauline leaves 0. Racemes 2–10 (–22) -flowered, ebracteate, not or slightly elongated in fruit; rachis not or slightly flexuous, glabrous or pubescent as stem. Fruiting pedicels divaricate-ascending to ascending (not decurrent basally), straight, (0.7–) 1.5–10 (–25) mm, usually glabrous, rarely sparsely pubescent, trichomes predominantly simple (0.2–0.7 mm), sometimes 2–4 (or 5) -rayed, (0.1–0.5 mm). Flowers: sepals ovate, 2–3 mm, usually sparsely pubescent, rarely glabrous, (trichomes simple and short-stalked, 2–4-rayed); petals pale-yellow (sometimes fading white), oblanceolate to obovate, 2–5 × 1–1.7 (–2) mm; anthers ovate or oblong, 0.4–0.6 mm. Fruits ovoid or ovatelanceolate, plane (not curved), flattened, (2.5–) 3–6 (–8) × 2–3 mm; valves pubescent or puberulent, trichomes simple and short-stalked, 2–5-rayed, 0.1–0.4 mm; ovules 4–12 per ovary; style 0.3–0.6 (–1) mm. Seeds oblong to ovoid, 1.2–2 (–2.6) × 0.9–1.2 (–1.4) mm. 2n = 36.


Phenology: Flowering Jun–Aug.
Habitat: Rock outcrops and talus, rocky knolls, alpine ridges
Elevation: 800-3700 m

Distribution

V7 401-distribution-map.gif

Alta., B.C., Alaska, Calif., Idaho, Nev., Oreg., Utah, Wash., Wyo.

Discussion

Draba densifolia is one of the most highly variable North American members of the genus, as evidenced by the extensive synonymy. The species is an apomict (G. A. Mulligan 1976) and it occupies tremendous geographic, edaphic, and altitudinal ranges. In forms corresponding to the type of D. sphaerula, the flowering stems are 2-flowered, and the stems, leaves, and fruiting pedicels are to 5, 1.5, and 0.6 mm, respectively. By contrast, forms comparable to the type of D. caeruleomontana var. piperi have stems, leaves, and fruiting pedicels to 17, 1.3, and 2.5 cm, respectively. Between these remarkable extremes fall all of the other populations of the species.

Draba densifolia is sometimes confused with D. paysonii, which it resembles, in being a cespitose, scapose, and densely pulvinate perennial having narrowly linear or oblong to linear-oblanceolate leaf blades with strongly ciliate margins. It is easily distinguished from the latter by having glabrous adaxial leaf blade surfaces, only sparsely pubescent abaxial surfaces, and often glabrous fruiting pedicels and rachises.

Selected References

None.

Lower Taxa

None.

"elongated" is not a number."thick" is not a number."dm" is not declared as a valid unit of measurement for this property."dm" is not declared as a valid unit of measurement for this property.

... more about "Draba densifolia"
oblong +  and ovate +
0.04 cm0.4 mm <br />4.0e-4 m <br /> (0.06 cm0.6 mm <br />6.0e-4 m <br />) +
Ihsan A. Al-Shehbaz +, Michael D. Windham +  and Reidar Elven +
Nuttall in J. Torrey and A. Gray +
appendaged +  and unappendaged +
0.9 cm9 mm <br />0.009 m <br /> (1.4 cm14 mm <br />0.014 m <br />) +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
not +  and auriculate +
0.3 cm3 mm <br />0.003 m <br /> (0.9 cm9 mm <br />0.009 m <br />) +
lobed +, toothed +  and entire +
linear;oblong or oblanceolate-linear +
much smaller +  and reduced +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
urceolate +, campanulate +  and tubular +
scapose +  and branched +
sessile +  and petiolate +
well-developed +
distinct +
well-differentiated +
emarginate +  and entire +
Alta. +, B.C. +, Alaska +, Calif. +, Idaho +, Nev. +, Oreg. +, Utah +, Wash. +  and Wyo. +
800-3700 m +
straight +  and curved +
not winged +  and unappendaged +
connate +  and distinct +
not basally +  and dilated +
actinomorphic +
orange;purple;orange;purple;pink;white;yellow +
spreading +, ascending +  and erect +
linear +, oblanceolate +, spatulate +, obovate +, subsaccate +, not saccate +, oblong +  and ovate +
latiseptate +, twisted +  and sessile +
0.6 cm6 mm <br />0.006 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
indehiscent +  and dehiscent +
0.3 cm3 mm <br />0.003 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
flattened +, plane +, ovatelanceolate +  and ovoid +
0.2 cm2 mm <br />0.002 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
Rock outcrops and talus, rocky knolls, alpine ridges +
sessile +  and petiolate +
alternate +, not +  and rosulate +
cauline +  and basal +
connate +  and distinct +
decurrent +
pinnatifid +
tenuinucellate +, crassinucellate +  and bitegmic +
campylotropous +  and anatropous +
0.17 cm1.7 mm <br />0.0017 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
pale-yellow +
rudimentary +
oblanceolate +  and obovate +
0.1 cm1 mm <br />0.001 m <br /> (0.17 cm1.7 mm <br />0.0017 m <br />) +
Flowering Jun–Aug. +
trinucleate +  and 3(-11)-colpate +
in J. Torrey and A. Gray, Fl. N. Amer. +
ebracteate;2-10(-22)-flowered +
ascending +  and erect +
slender +
pubescent +  and glabrous +
mucilaginous +
0.2 cm2 mm <br />0.002 m <br /> (0.26 cm2.6 mm <br />0.0026 m <br />) +
0.12 cm1.2 mm <br />0.0012 m <br /> (0.14 cm1.4 mm <br />0.0014 m <br />) +
not +  and mucilaginous +
white +, black +, brown +  and yellow +
0.12 cm1.2 mm <br />0.0012 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
oblong +  and ovoid +
0.09 cm0.9 mm <br />9.0e-4 m <br /> (0.12 cm1.2 mm <br />0.0012 m <br />) +
persistent +  and caducous +
distinct +
spreading +, ascending +  and erect +
glabrous +  and pubescent +
0.2 cm2 mm <br />0.002 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
reduced +
latiseptate +, twisted +  and sessile +
indehiscent +  and dehiscent +
terete +, plane +, subglobose +, ovoid +, suborbicular +, linear +, oblong +, elliptic +, lanceolate +  and ovate +
latiseptate +, twisted +  and sessile +
indehiscent +  and dehiscent +
terete +, plane +, subglobose +, ovoid +, suborbicular +, linear +, oblong +, elliptic +, lanceolate +  and ovate +
Illustrated +  and Endemic +
tetradynamous +
procumbent +  and decumbent +
prostrate +  and decumbent +
usually erect +  and ascending +
pubescent +  and glabrous +
subsessile +, sessile +  and petiolate +
0.06 cm0.6 mm <br />6.0e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
persistent +
distinct +
0.03 cm0.3 mm <br />3.0e-4 m <br /> (0.06 cm0.6 mm <br />6.0e-4 m <br />) +
pubescent +  and glabrous +
0.01 cm0.1 mm <br />1.0e-4 m <br /> (0.03 cm0.3 mm <br />3.0e-4 m <br />) +
Draba caeruleomontana +, Draba caeruleomontana var. piperi +, Draba glacialis var. pectinata +, Draba globosa var. sphaerula +, Draba mulfordiae +, Draba nelsonii +, Draba oligosperma var. pectinata +, Draba pectinata +  and Draba sphaerula +
Draba densifolia +
species +
short-stalked +  and simple +
cruciform +  and forked +
0.01 cm0.1 mm <br />1.0e-4 m <br /> (0.04 cm0.4 mm <br />4.0e-4 m <br />) +
coiled +, 1-7-veined +  and veined +
puberulent +  and pubescent +
anastomosing +
scapose +  and not +
1 cm10 mm <br />0.01 m <br /> (2.5 cm25 mm <br />0.025 m <br />) +
divaricate-ascending +  and ascending +
pubescent +  and glabrous +
0.15 cm1.5 mm <br />0.0015 m <br /> (1 cm10 mm <br />0.01 m <br />) +
rhizomatous +, taprooted +, scapose +  and not +
perennial +, biennial +  and annual +
subshrub +  and herb +
aquatics +  and terrestrial +
glabrous +  and pubescent +