Draba fladnizensis

Wulfen in N. J. Jacquin

in N. J. Jacquin, Misc. Austriac. 1: 147, plate 17, fig. 1. 1778.

Synonyms: Draba fladnizensis var. pattersonii (O. E. Schulz) Rollins Draba pattersonii Hartman Draba pattersonii var. hirticaulis O. E. Schulz Draba wahlenbergii
Treatment appears in FNA Volume 7. Treatment on page 307. Mentioned on page 270, 275, 281, 303, 316, 320.

Perennials; (sometimes cespitose); caudex simple or branched (with persistent leaf-bases); usually scapose. Stems unbranched, (0.2–) 0.3–1 (–1.3) dm, glabrous. Basal leaves rosulate; petiole (obscure), margin ciliate, (trichomes simple or 2-rayed, 0.25–0.6 mm); blade linear to oblanceolate or narrowly obovate, (0.3–) 0.4–1.2 (–1.6) cm × 1–3 (–4) mm, margins usually entire, rarely toothed, surfaces abaxially pubescent or glabrous, trichomes simple, sometimes with fewer, short-stalked, 2-rayed ones, (midvein prominent), adaxially often glabrous. Cauline leaves 0–2; sessile; blade oblong to ovate, margins entire, (ciliate). Racemes (2 or) 3–11 (–14) -flowered, usually ebracteate, rarely proximalmost flowers bracteate, elongated in fruit; rachis not flexuous, glabrous. Fruiting pedicels divaricate-ascending, often straight, (1–) 2–5 (–6) mm, glabrous. Flowers: sepals (green or purplish), ovate, 1.2–2.2 mm, glabrous or pubescent, (trichomes simple); petals white, spatulate, 2–2.5 × 0.8–1.5 mm; anthers ovate, 0.2–0.25 mm. Fruits elliptic-lanceolate to oblong, plane, flattened, 3–8 (–9) × 1.5–2 mm; valves glabrous; ovules 12–24 per ovary; style 0.05–0.2 (–0.3) mm. Seeds oblong to elliptic, 0.8–1 × 0.5–0.6 mm. 2n = 16.


Phenology: Flowering Jun–Aug.
Habitat: Rock outcrops and talus, alpine meadows, sandy gravel
Elevation: 0-1400 m at higher latitudes, 3000-3800 m at lower latitudes

Distribution

V7 403-distribution-map.gif

Greenland, B.C., N.W.T., Nunavut, Que., Yukon, Alaska, Colo., Utah, Wyo., c, s Europe, Asia, circumpolar and high alpine areas

Discussion

Rollins reduced Draba pattersonii to a variety of D. fladnizensis and separated the two primarily on plant size and minor differences in fruit shape. Examination of D. fladnizensis specimens collected throughout Europe and North America reveals that the alleged differences between the two taxa are artificial. The type material of D. pattersonii, which was collected in Colorado, is a mixture of plants highly variable in their type of indumentum. The specimens have no flowers, but the habit, fruits, and leaves are nearly indistinguishable from those of D. fladnizensis from higher latitudes.

In the absence of flowers, the white-flowered Draba fladnizensis (2n = 16) is often confused with the yellow-flowered D. crassifolia (2n = 40). The latter is an annual or short-lived perennial that rarely forms a well-developed caudex, whereas D. fladnizensis almost always produces a distinct caudex. Although most individuals of both species are scapose, they occasionally produce one or two cauline leaves. The cauline leaves are usually glabrous in D. crassifolia and ciliate in D. fladnizensis; in the latter, the distalmost cauline leaf usually subtends the proximalmost flower. Finally, the seeds in D. fladnizensis are slightly larger (0.8–0.1 × 0.5–0.6 versus 0.7–0.8 × 0.4–0.5 mm) than those of D. crassifolia, though the reliability of this distinction needs to be examined in greater detail. N. H. Holmgren (2005b) reported D. fladnizensis from central Nevada, but we have not seen any material of the species from that state.

Selected References

None.

Lower Taxa

None.

"elongated" is not a number."elongated" is not a number."thick" is not a number."dm" is not declared as a valid unit of measurement for this property.

... more about "Draba fladnizensis"
0.02 cm0.2 mm <br />2.0e-4 m <br /> (0.025 cm0.25 mm <br />2.5e-4 m <br />) +
Ihsan A. Al-Shehbaz +, Michael D. Windham +  and Reidar Elven +
Wulfen in N. J. Jacquin +
sessile +  and petiolate +
appendaged +  and unappendaged +
1.2 cm12 mm <br />0.012 m <br /> (1.6 cm16 mm <br />0.016 m <br />) +
0.3 cm3 mm <br />0.003 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
0.4 cm4 mm <br />0.004 m <br /> (1.2 cm12 mm <br />0.012 m <br />) +
lobed +, toothed +  and entire +
oblong;ovate +
much smaller +  and reduced +
1mm;3mm +
urceolate +, campanulate +  and tubular +
scapose +, branched +  and simple +
well-developed +
distinct +
well-differentiated +
emarginate +  and entire +
Greenland +, B.C. +, N.W.T. +, Nunavut +, Que. +, Yukon +, Alaska +, Colo. +, Utah +, Wyo. +, c +, s Europe +, Asia +  and circumpolar and high alpine areas +
0-1400 m at higher latitudes, 3000-3800 m at lower latitudes +
straight +  and curved +
not winged +  and unappendaged +
connate +  and distinct +
not basally +  and dilated +
orange;purple;orange;purple;pink;white;yellow +
spreading +, ascending +  and erect +
proximalmost +
pubescent +  and glabrous +
0.12 cm1.2 mm <br />0.0012 m <br /> (0.22 cm2.2 mm <br />0.0022 m <br />) +
latiseptate +, twisted +  and sessile +
0.8 cm8 mm <br />0.008 m <br /> (0.9 cm9 mm <br />0.009 m <br />) +
indehiscent +  and dehiscent +
flattened +, plane +, elliptic-lanceolate +  and oblong +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
Rock outcrops and talus, alpine meadows, sandy gravel +
sessile +  and petiolate +
alternate +, not +  and rosulate +
cauline +  and basal +
connate +  and distinct +
decurrent +
tenuinucellate +, crassinucellate +  and bitegmic +
campylotropous +  and anatropous +
0.2 cm2 mm <br />0.002 m <br /> (0.25 cm2.5 mm <br />0.0025 m <br />) +
rudimentary +
0.08 cm0.8 mm <br />8.0e-4 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
Flowering Jun–Aug. +
trinucleate +  and 3(-11)-colpate +
in N. J. Jacquin, Misc. Austriac. +
ebracteate;3-11(-14)-flowered +
not +  and elongated +
ascending +  and erect +
slender +
0.5 cm5 mm <br />0.005 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
straight +  and not flexuous +
divaricate-ascending +
mucilaginous +
not +  and mucilaginous +
White (?) +, Black (?) +, Brown (?) +  and Yellow (?) +
0.08 cm0.8 mm <br />8.0e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
oblong +  and elliptic +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.06 cm0.6 mm <br />6.0e-4 m <br />) +
persistent +  and caducous +
distinct +
spreading +, ascending +  and erect +
reduced +
latiseptate +, twisted +  and sessile +
indehiscent +  and dehiscent +
terete +, plane +, subglobose +, ovoid +, suborbicular +, linear +, oblong +, elliptic +, lanceolate +  and ovate +
latiseptate +, twisted +  and sessile +
indehiscent +  and dehiscent +
terete +, plane +, subglobose +, ovoid +, suborbicular +, linear +, oblong +, elliptic +, lanceolate +  and ovate +
tetradynamous +
procumbent +  and decumbent +
prostrate +  and decumbent +
usually erect +  and ascending +
subsessile +, sessile +  and petiolate +
0.02 cm0.2 mm <br />2.0e-4 m <br /> (0.03 cm0.3 mm <br />3.0e-4 m <br />) +
persistent +
distinct +
0.005 cm0.05 mm <br />5.0e-5 m <br /> (0.02 cm0.2 mm <br />2.0e-4 m <br />) +
glabrous +  and pubescent +
Draba fladnizensis var. pattersonii +, Draba pattersonii +, Draba pattersonii var. hirticaulis +  and Draba wahlenbergii +
Draba fladnizensis +
species +
cruciform +  and forked +
coiled +, 1-7-veined +  and veined +
anastomosing +
scapose +  and not +
pubescent +  and glabrous +
rhizomatous +, taprooted +, scapose +  and not +
perennial +, biennial +  and annual +
subshrub +  and herb +
aquatics +  and terrestrial +
glabrous +  and pubescent +