Draba glabella

Pursh

Fl. Amer. Sept. 2: 434. 1813.

Synonyms: Draba arabisans var. canadensis (Brunet) Fernald & Knowlton Draba arabisans var. orthocarpa Fernald & Knowlton Draba canadensis de Candolle Draba canadensis var. pycnosperma (Fernald & Knowlton) O. E. Schulz Draba daurica Linnaeus Draba glabella var. megasperma (Fernald & Knowlton) Fernald Draba glabella var. orthocarpa (Fernald & Knowlton) Fernald Draba glabella var. pycnosperma (Fernald & Knowlton) G. A. Mulligan Draba henneana Fernald & Knowlton Draba hirta var. laurentiana (Fernald) B. Boivin Draba hirta var. pycnosperma (Fernald & Knowlton) B. Boivin Draba laurentiana Draba megasperma Draba norvegica var. pleiophylla Fernald Draba pycnosperma Draba sornborgeri
Treatment appears in FNA Volume 7. Treatment on page 307. Mentioned on page 275, 277, 291, 295, 296, 308.

Perennials; (sometimes cespitose); caudex simple or branched; not scapose. Stems branched or unbranched, (0.4–) 1–3.5 (–4.7) dm, often pubescent throughout (sometimes sparsely so distally), sometimes glabrous, trichomes simple and 2-rayed, (non-crisped), 0.2–1 mm, or subsessile, stellate-pectinate, and 3–8-rayed, 0.1–0.3 mm. Basal leaves rosulate; petiolate; petiole ciliate, (trichomes simple, 0.1–0.8 mm); blade oblanceolate to spatulate or linear-oblanceolate, (0.6–) 1–3.5 (–5) cm × 2–8 (–10) mm, margins dentate or denticulate, (pubescent as petiole), surfaces pubescent with non-crisped, minutely stalked, 4–8 (–12) -rayed, stellate-pectinate trichomes, 0.15–0.6 mm, (midvein obscure abaxially), adaxially sometimes also with simple trichomes, or glabrous. Cauline leaves 2–17 (–25); sessile; blade ovate to oblong, margins dentate to subentire, surfaces often pubescent as basal, or predominantly with simple trichomes adaxially. Racemes (5–) 8–26 (–34) -flowered, ebracteate or proximalmost 1 or 2 flowers bracteate, elongated in fruit; rachis not flexuous, usually glabrous, rarely pubescent as stem. Fruiting pedicels divaricate-ascending to suberect, straight, (1–) 3–10 (–16) mm, glabrous or pubescent as stem. Flowers: sepals oblong, 2–3.5 mm, pubescent, (trichomes simple and short-stalked, 2–4-rayed); petals white, broadly obovate, 4–5.5 × 1.5–3 mm; anthers ovate, 0.3–0.5 mm. Fruits oblong to ovate or ovoid to lanceolate or linear-lanceolate, usually plane, rarely slightly twisted, flattened or inflated, (3–) 5–12 (–16) × 2–3.5 mm; valves glabrous or pubescent, trichomes simple or 2–4-rayed, 0.05–0.2 (–0.4) mm; ovules (20–) 24–36 per ovary; style 0.05–0.2 (–0.5) mm. Seeds oblong, 0.9–1.1 × 0.5–0.7 mm. 2n = 64, 80.


Phenology: Flowering May–Aug.
Habitat: Rock outcrops, talus, rocky ridges and knolls, meadows, tundra, gravelly beaches sandy river margins, disturbed soils
Elevation: 0-1400 m

Distribution

V7 404-distribution-map.gif

Greenland, B.C., Man., N.B., Nfld. and Labr., N.W.T., N.S., Nunavut, Ont., Que., Yukon, Alaska, Maine, Vt., Wis., n Europe (n Russia), e Asia (Russian Far East), e Asia (Siberia)

Discussion

Draba glabella was reported by J. V. Freudenstein and J. K. Marr (1986) from Michigan, but that record likely was based on plants of D. arabisans, a highly variable species that occurs in that state.

Draba glabella is extremely variable in indumentum, number of cauline leaves, fruiting pedicel length, fruit shape and size, style length, and seed number. M. L. Fernald (1934) divided it into five species and three varieties; R. C. Rollins (1993) recognized three species. Of these, G. A. Mulligan (1970, 1976) reduced D. laurentiana to synonymy of D. glabella and treated D. pycnosperma as a variety. Of all the segregates of D. glabella, var. pycnosperma might merit recognition. It is restricted to northwestern Newfoundland and northeastern Quebec, where var. glabella also grows. The main difference between the two varieties is the presence in var. pycnosperma of plump (versus flattened), ovoid to oblong fruits. Draba sornborgeri, recognized by Rollins as a distinct species, is merely a glabrescent form of D. glabella. Because of the tremendous morphological variability, wide distribution, extensive synonymy, and different chromosome numbers, D. glabella will require extensive molecular, cytological, and morphological studies to properly delimit the species and any potential infraspecific taxa.

Some forms of Draba glabella approach both D. borealis and D. praealta, but these can be distinguished by examining the trichomes on the abaxial surfaces of basal leaf blades. In D. glabella, these trichomes are minutely stalked or subsessile and have branched rays. In the other two species, the trichomes have long stalks and the rays are always unbranched.

The Linnaean name Draba hirta was applied to this species previously, and still is in Russia. The name is not typified, and the material at LINN is in bad condition and probably belongs to two species, D. glabella and D. norvegica, as recognized here. A typification of D. hirta may necessitate its re-introduction for this species.

Selected References

None.

Lower Taxa

None.

"elongated" is not a number."thick" is not a number.

... more about "Draba glabella"
0.03 cm0.3 mm <br />3.0e-4 m <br /> (0.05 cm0.5 mm <br />5.0e-4 m <br />) +
Ihsan A. Al-Shehbaz +, Michael D. Windham +  and Reidar Elven +
appendaged +  and unappendaged +
3.5 cm35 mm <br />0.035 m <br /> (5 cm50 mm <br />0.05 m <br />) +
0.8 cm8 mm <br />0.008 m <br /> (1 cm10 mm <br />0.01 m <br />) +
not +  and auriculate +
1 cm10 mm <br />0.01 m <br /> (3.5 cm35 mm <br />0.035 m <br />) +
lobed +, toothed +  and entire +
ovate;oblong +
much smaller +  and reduced +
0.2 cm2 mm <br />0.002 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
urceolate +, campanulate +  and tubular +
not scapose +, branched +  and simple +
well-developed +
distinct +
well-differentiated +
emarginate +  and entire +
Greenland +, B.C. +, Man. +, N.B. +, Nfld. and Labr. +, N.W.T. +, N.S. +, Nunavut +, Ont. +, Que. +, Yukon +, Alaska +, Maine +, Vt. +, Wis. +, n Europe (n Russia) +, e Asia (Russian Far East) +  and e Asia (Siberia) +
0-1400 m +
straight +  and curved +
not winged +  and unappendaged +
connate +  and distinct +
not basally +  and dilated +
actinomorphic +
orange;purple;orange;purple;pink;white;yellow +
spreading +, ascending +  and erect +
linear +, oblanceolate +, spatulate +, obovate +, subsaccate +, not saccate +, oblong +  and ovate +
1.2 cm12 mm <br />0.012 m <br /> (1.6 cm16 mm <br />0.016 m <br />) +
indehiscent +  and dehiscent +
0.5 cm5 mm <br />0.005 m <br /> (1.2 cm12 mm <br />0.012 m <br />) +
inflated +, flattened +, plane +, oblong +  and ovate or ovoid +
0.2 cm2 mm <br />0.002 m <br /> (0.35 cm3.5 mm <br />0.0035 m <br />) +
Rock outcrops, talus, rocky ridges and knolls, meadows, tundra, gravelly beaches sandy river margins, disturbed soils +
sessile +  and petiolate +
alternate +, not +  and rosulate +
cauline +  and basal +
connate +  and distinct +
decurrent +
dentate +  and entire +
dentate;subentire +
tenuinucellate +, crassinucellate +  and bitegmic +
campylotropous +  and anatropous +
4mm;5.5mm +
rudimentary +
obovate +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
Flowering May–Aug. +
trinucleate +  and 3(-11)-colpate +
Fl. Amer. Sept. +
bracteate;ebracteate;(5-)8-26(-34)-flowered +
ascending +  and erect +
proximalmost +
slender +
not flexuous +
pubescent +  and glabrous +
mucilaginous +
not +  and mucilaginous +
white +, black +, brown +  and yellow +
0.09 cm0.9 mm <br />9.0e-4 m <br /> (0.11 cm1.1 mm <br />0.0011 m <br />) +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.07 cm0.7 mm <br />7.0e-4 m <br />) +
persistent +  and caducous +
distinct +
spreading +, ascending +  and erect +
0.2 cm2 mm <br />0.002 m <br /> (0.35 cm3.5 mm <br />0.0035 m <br />) +
reduced +
latiseptate +, twisted +  and sessile +
indehiscent +  and dehiscent +
terete +, plane +, subglobose +, ovoid +, suborbicular +, linear +, oblong +, elliptic +, lanceolate +  and ovate +
latiseptate +, twisted +  and sessile +
indehiscent +  and dehiscent +
terete +, plane +, subglobose +, ovoid +, suborbicular +, linear +, oblong +, elliptic +, lanceolate +  and ovate +
tetradynamous +
unbranched +  and branched +
1 cm10 mm <br />0.01 m <br /> (1.6 cm16 mm <br />0.016 m <br />) +
straight +
procumbent +  and decumbent +
prostrate +  and decumbent +
divaricate-ascending +  and suberect +
pubescent +  and glabrous +
0.3 cm3 mm <br />0.003 m <br /> (1 cm10 mm <br />0.01 m <br />) +
subsessile +, sessile +  and petiolate +
0.02 cm0.2 mm <br />2.0e-4 m <br /> (0.05 cm0.5 mm <br />5.0e-4 m <br />) +
persistent +
distinct +
0.005 cm0.05 mm <br />5.0e-5 m <br /> (0.02 cm0.2 mm <br />2.0e-4 m <br />) +
pubescent +  and glabrous +
0.015 cm0.15 mm <br />1.5e-4 m <br /> (0.06 cm0.6 mm <br />6.0e-4 m <br />) +
Draba arabisans var. canadensis +, Draba arabisans var. orthocarpa +, Draba canadensis +, Draba canadensis var. pycnosperma +, Draba daurica +, Draba glabella var. megasperma +, Draba glabella var. orthocarpa +, Draba glabella var. pycnosperma +, Draba henneana +, Draba hirta var. laurentiana +, Draba hirta var. pycnosperma +, Draba laurentiana +, Draba megasperma +, Draba norvegica var. pleiophylla +, Draba pycnosperma +  and Draba sornborgeri +
Draba glabella +
species +
2-4-rayed +, simple +, stalked +  and subsessile +
0.02 cm0.2 mm <br />2.0e-4 m <br /> (0.04 cm0.4 mm <br />4.0e-4 m <br />) +
stellate-pectinate +  and non-crisped +
0.005 cm0.05 mm <br />5.0e-5 m <br /> (0.02 cm0.2 mm <br />2.0e-4 m <br />) +
coiled +, 1-7-veined +  and veined +
pubescent +  and glabrous +
anastomosing +
scapose +  and not +
pubescent +  and glabrous +
rhizomatous +, taprooted +, scapose +  and not +
perennial +, biennial +  and annual +
subshrub +  and herb +
aquatics +  and terrestrial +
glabrous +  and pubescent +